Linyphiidae Blackwall, 1859

Family |   ACCEPTED
N.B.: for comments on relationships, see Michalik & Hormiga, 2010: 7; re-synonymization of Pimoidae (Murphy & Roberts, 2015: ix) not sufficiently justified and is not accepted here; Sinopimoidae Li & Wunderlich, 2008 [urn:lsid:nmbe.ch:spiderfam:0112] synonymyized with Linyphiidae by Dimitrov et al., 2017: 245; resurrection of Sinopimoidae by Wunderlich, 2020b: 20 is not accepted here, because Dimitrov et al., 2017 was not taken into account; re-elevation of Sinopimoidae to family level by Eskov & Marusik, 2023: 343 not accepted here (by majority vote of the WSC boards); the current status of Sinopimoidae is also maintained for reasons of stability and we wait for a probable confirmation or rejection of this resurrection; Tian et al., 2022 show five subfamilies: Linyphiinae (incl. Mynogleninae  Lehtinen, 1967: 250 and Dubiaraneinae Millidge, 1993c: 145), Erigoninae Emerton, 1882: 7, Micronetinae Hull, 1920a: 8, Ipainae Saaristo, 2007b: 34 and Stemonyphantinae Wunderlich, 1986: 120; da Silva-Moreira, Kulkarni & Hormiga, 2025: 60-62 list resp. newly circumscribe six subfamilies: Erigoninae, Lepthyphantinae Simon, 1929: 562, Linyphiinae (syn. nov.: Dubiaraneinae), Mynogleninae, Pocobletinae Silva-Moreira & Hormiga, 2025: 61 (new subfamily) and Stemonyphantinae, state Ipainae and Micronetinae to be polyphyletic, leave several linyphiid clades unranked and also several genera as incertae sedis.
Gen. Megalepthyphantes Wunderlich, 1994 urn:lsid:nmbe.ch:spidergen:01086
type Linyphia nebulosa Sundevall, 1830; gender masculine
N.B.: contains the nebulosus-group sensu Wunderlich, 1977b (Wunderlich, 1994b: 169)

Transferred to other genera:

Megalepthyphantes globularis Tanasevitch, 2011 -- see Plesiophantes

In synonymy:

Megalepthyphantes autumnalis (Hentz, 1850, T from Linyphia) = Megalepthyphantes nebulosus (Sundevall, 1830) (Emerton, 1882: 69, sub Bathyphantes)

Megalepthyphantes circumflexus (C. L. Koch, 1845) = Megalepthyphantes nebulosus (Sundevall, 1830) (Thorell, 1870a: 54, sub Linyphia)

Megalepthyphantes furcula (C. L. Koch, 1837) = Megalepthyphantes nebulosus (Sundevall, 1830) (Thorell, 1870a: 54, sub Linyphia)

Megalepthyphantes klingelbachi (Wunderlich, 1977) = Megalepthyphantes collinus (L. Koch, 1872) (Saaristo & Tanasevitch, 1993: 60, sub Lepthyphantes occidentalis)

Megalepthyphantes occidentalis (Machado, 1949, T from Lepthyphantes) = Megalepthyphantes collinus (L. Koch, 1872) (Saaristo, 1997a: 259)

Megalepthyphantes vivax (Blackwall, 1841) = Megalepthyphantes nebulosus (Sundevall, 1830) (Thorell, 1870a: 54, sub Linyphia)

Megalepthyphantes xinjiangensis (Hu & Wu, 1989) = Megalepthyphantes kronebergi (Tanasevitch, 1989) (Tanasevitch, 1992: 48, sub Lepthyphantes)

Megalepthyphantes auresensis Bosmans, 2006 | | Algeria urn:lsid:nmbe.ch:spidersp:039914
Megalepthyphantes auresensis Bosmans, 2006b: 181, f. 18-24 (Dmf) Original description
Megalepthyphantes bkheitae (Bosmans & Bouragba, 1992) | | Algeria urn:lsid:nmbe.ch:spidersp:011581
Lepthyphantes bkheitae Bosmans & Bouragba, 1992: 251, f. 11-16 (Dmf) Original description
Megalepthyphantes bkheitae Saaristo & Tanasevitch, 2000: 264 (T from Lepthyphantes)
Megalepthyphantes brignolii Tanasevitch, 2014 | | Morocco urn:lsid:nmbe.ch:spidersp:047248
Megalepthyphantes brignolii Tanasevitch, 2014c: 285, f. 25-28 (Dmf) Original description
Megalepthyphantes camelus (Tanasevitch, 1990) | | Iran, Azerbaijan urn:lsid:nmbe.ch:spidersp:011582
Lepthyphantes camelus Tanasevitch, 1990: 19, f. 6.1 (Dm) Original description
Megalepthyphantes camelus Saaristo & Tanasevitch, 2000: 264 (T from Lepthyphantes)
Megalepthyphantes camelus Tanasevitch, 2008b: 481, f. 29-31 (Df)
Megalepthyphantes camelus Tanasevitch, 2009a: 398, f. 38-42 (mf)
Megalepthyphantes camelus Zamani & Marusik, 2018c: 71, f. 2D, 4A-C (m)
Megalepthyphantes collinus (L. Koch, 1872) | | Europe, Turkey urn:lsid:nmbe.ch:spidersp:011583
Linyphia collina L. Koch, 1872c: 134, pl. 1, f. 9-12 (Dmf) Original description
Bathyphantes collinus Förster & Bertkau, 1883: 273
Lepthyphantes collinus Simon, 1884a: 275, f. 26-27 (mf)
Lephthyphantes collinus Chyzer & Kulczyński, 1894: 66, pl. 2, f. 31 (mf)
Lepthyphantes collinus Simon, 1894a: 688, f. 772 (m)
Lepthyphantes collinus Bösenberg, 1901: 73, pl. 6, f. 75 (mf)
Lepthyphantes collinus Simon, 1929: 578, 729, f. 863-864 (mf)
Lepthyphantes collinus occidentalis Machado, 1949: 37, f. 17 (Dm) [urn:lsid:nmbe.ch:spidersp:056265]
Lepthyphantes collinus Wiehle, 1956: 168, f. 283-285 (mf)
Lepthyphantes collinus Tystshenko, 1971: 218, f. 659, 678 (mf)
Lepthyphantes collinus Staręga, 1972: 66, f. 1-2, photo 1 (f)
Lepthyphantes klingelbachi Wunderlich, 1977b: 57, f. 1-2 (Dm) [urn:lsid:nmbe.ch:spidersp:056264]
Lepthyphantes cf. collinus Deeleman-Reinhold, 1984: 43, f. 15-18 (f)
Lepthyphantes collinus Bosmans, 1985a: 142, f. 1d-f (f)
Lepthyphantes collinus Tanasevitch, 1990: 96, f. 25.7, 26.17 (mf)
Lepthyphantes collinus Heimer & Nentwig, 1991: 180, f. 488 (mf)
Lepthyphantes occidentalis Saaristo & Tanasevitch, 1993: 60 (elevated from subspecies of Lepthyphantes collinus, S of Lepthyphantes klingelbachi)
Megalepthyphantes collinus Saaristo, 1997a: 259, f. 5-8 (mf, T from Lepthyphantes, S of Lepthyphantes occidentalis)
Megalepthyphantes collinus Höfer et al., 2019: 68, f. 15a-b (m)
Megalepthyphantes near collinus Berry, 2020: 6, f. 1-3 (m)
Megalepthyphantes sp. near collinus Parker, 2021b: 10, f. 1-4 (mf)
Megalepthyphantes sp. near collinus Cann, 2022: 1, f. 1-2 (f)
Megalepthyphantes gongshanensis Irfan, Zhang & Peng, 2022 | | China urn:lsid:nmbe.ch:spidersp:057083
Megalepthyphantes gongshanensis Irfan, Zhang & Peng, 2022: 162, f. 184A-D, 185A-D, 186A-D (Dmf) Original description
Megalepthyphantes hellinckxorum Bosmans, 2006 | | Algeria urn:lsid:nmbe.ch:spidersp:039915
Megalepthyphantes hellinckxorum Bosmans, 2006b: 183, f. 25-31 (Dmf) Original description
Megalepthyphantes kandahar Tanasevitch, 2009 | | Afghanistan urn:lsid:nmbe.ch:spidersp:042908
Megalepthyphantes kandahar Tanasevitch, 2009b: 423, f. 1-6 (Dmf) Original description
Megalepthyphantes kronebergi (Tanasevitch, 1989) | | Iran, Kazakhstan to China urn:lsid:nmbe.ch:spidersp:011584
Lepthyphantes xinjiangensis Hu & Wu, 1989: 150, f. 124.1-6 (Dmf) [urn:lsid:nmbe.ch:spidersp:056266]
Lepthyphantes kronebergi Tanasevitch, 1989a: 101, f. 41-45 (Dmf) Original description
Lepthyphantes kronebergi Tanasevitch, 1992: 48 (S of Lepthyphantes xinjiangensis)
Lepthyphantes kronebergi Song, Zhu & Chen, 1999: 182, f. 102K-L, T (mf)
Megalepthyphantes kronebergi Saaristo & Tanasevitch, 2000: 264 (T from Lepthyphantes)
Megalepthyphantes kuhitangensis (Tanasevitch, 1989) | | Israel, Iran, Central Asia, China urn:lsid:nmbe.ch:spidersp:011204
Lepthyphantes kuhitangensis Tanasevitch, 1989a: 102, f. 38-40 (Df) Original description
Lepthyphantes kuhitangensis Song, Zhu & Chen, 1999: 182, f. 102M-N (f)
Megalepthyphantes kuhitangensis Tanasevitch, 2008b: 481, f. 32 (f, T from Lepthyphantes)
Megalepthyphantes kuhitangensis Tanasevitch, 2009a: 400, f. 43-46, 50-52 (f, Dm)
Megalepthyphantes kuhitangensis Zamani, Nadolny & Dolejš, 2022: 556, f. 2A-H (mf)
Megalepthyphantes lydiae Wunderlich, 1994 | | Greece urn:lsid:nmbe.ch:spidersp:011585
Megalepthyphantes lydiae Wunderlich, 1994b: 169, f. 1-5 (Dmf) Original description
Megalepthyphantes lydiae Tanasevitch, 2019i: 147, f. 1-13 (mf)
Megalepthyphantes minotaur Tanasevitch & Wunderlich, 2015 | | Greece (Crete) urn:lsid:nmbe.ch:spidersp:047736
?Megalepthyphantes minotaur Tanasevitch & Wunderlich, 2015: 453, f. 1-7 (Dm) Original description
Megalepthyphantes nebulosoides (Wunderlich, 1977) | | Iran, Central Asia, Nepal urn:lsid:nmbe.ch:spidersp:011586
Lepthyphantes nebulosoides Wunderlich, 1977b: 59, f. 9-12 (Dmf) Original description
Lepthyphantes nebulosoides Tanasevitch, 1989a: 104, f. 52-54 (mf)
Megalepthyphantes nebulosoides Saaristo, 1997a: 257 (T from Lepthyphantes)
Megalepthyphantes nebulosoides Tanasevitch & Saaristo, 2006: 13, f. 1-4 (f)
Megalepthyphantes nebulosus (Sundevall, 1830) | | North America, Europe, Turkey, Caucasus, Russia (Europe to Far East), Iraq, Kazakhstan urn:lsid:nmbe.ch:spidersp:011587 TYPE SPECIES
Linyphia nebulosa Sundevall, 1830: 218 (Dmf) Original description
Linyphia furcula C. L. Koch, 1837b: 10 (D) [urn:lsid:nmbe.ch:spidersp:065334]
Linyphia vivax Blackwall, 1841a: 657 (Dmf) [urn:lsid:nmbe.ch:spidersp:065335]
Linyphia furcula C. L. Koch, 1845: 116, f. 1040 (Df)
Linyphia circumflexa C. L. Koch, 1845: 128, f. 1050 (Dm) [urn:lsid:nmbe.ch:spidersp:065336]
Linyphia autumnalis Hentz, 1850: 30, pl. 4, f. 9 (Df) [urn:lsid:nmbe.ch:spidersp:065337]
Linyphia crypticola Westring, 1851: 37 (misidentified per Thorell, 1870a: 54)
Linyphia crypticola Westring, 1861: 113 (misidentified per Thorell, 1870a: 54)
Linyphia vivax Blackwall, 1864a: 221, pl. 16, f. 146 (mf)
Lepthyphantes crypticola Menge, 1866: 133, pl. 25, f. 54 (mf; misidentified per Thorell, 1870a: 54)
Linyphia circumflexa Ohlert, 1867: 45 (Df)
Linyphia nebulosa Thorell, 1870a: 54 (S of Linyphia circumflexa, L. furcula and L. vivax)
Bathyphantes nebulosus Emerton, 1882: 69, pl. 22, f. 1 (mf, S of Linyphia autumnalis)
Lepthyphantes nebulosus Simon, 1884a: 273, f. 23 (mf)
Linyphia nebulosus Keyserling, 1886b: 75, pl. 13, f. 174 (mf)
Lepthyphantes nebulosus Becker, 1896: 39, pl. 6, f. 2 (mf)
Lepthyphantes nebulosus Bösenberg, 1901: 72, pl. 6, f. 74 (mf)
Lepthyphantes nebulosus Smith, 1901b: 197, p. 109: f. h-k (mf)
Linyphia nebulosus Emerton, 1902: 143, f. 336-341 (mf)
Lepthyphantes nebulosus Bishop, 1925: 66, f. 2-3 (m)
Lepthyphantes nebulosus Spassky, 1925: 25, f. 18, 25 (mf)
Lepthyphantes nebulosus Simon, 1929: 578, 730, f. 861 (mf)
Lepthyphantes nebulosus Zorsch, 1937: 864, pl. 2, f. 16-23 (mf)
Lepthyphantes nebulosus Comstock, 1940: 394, f. 391 (f)
Lepthyphantes nebulosus Muma, 1943: 75, pl. XIII, f. 10 (f)
Lepthyphantes nebulosa Kaston, 1948: 128, f. 276-283 (mf)
Lepthyphantes nebulosus Locket & Millidge, 1953: 380, f. 226A, 227B, 228A (mf)
Lepthyphantes nebulosus Wiehle, 1956: 164, f. 277-282 (mf)
Lepthyphantes nebulosus Buchar, 1961: 89, f. 1E-F (f)
Lepthyphantes nebulosus Merrett, 1963b: 357, f. 9A-G (m)
Lepthyphantes nebulosus Azheganova, 1968: 64, f. 125, 131 (mf)
Lepthyphantes nebulosus Tystshenko, 1971: 216, f. 660, 676 (mf)
Lepthyphantes nebulosus Miller, 1971: 234, pl. XLV, f. 18-19 (mf)
Lepthyphantes nebulosus Wanless, 1971: 21, f. 1A, 5A (f)
Lepthyphantes nebulosus Wanless, 1973: 130, f. II.4, VII.10 (f)
Lepthyphantes nebulosus Palmgren, 1975: 54, f. 11.9-10 (mf)
Lepthyphantes nebulosus Wunderlich, 1977b: 59, f. 3-6 (mf)
Lepthyphantes nebulosus Millidge, 1984b: 248, f. 93 (f)
Lepthyphantes nebulosus Millidge, 1986: 58, f. 8
Lepthyphantes nebulosus Roberts, 1987: 148, f. 77a (mf)
Lepthyphantes nebulosus Hu & Wu, 1989: 147, f. 120.1-2 (f)
Lepthyphantes nebulosus Izmailova, 1989: 72, f. 51 (m)
Lepthyphantes nebulosus Tanasevitch, 1989a: 104, f. 55-56 (mf)
Lepthyphantes nebulosus Tanasevitch, 1990: 96, f. 25.6, 26.4 (mf)
Lepthyphantes nebulosus Heimer & Nentwig, 1991: 180, f. 487 (mf)
Lepthyphantes nebulosus Zhao, 1993: 156, f. 75a-b (f)
Megalepthyphantes nebulosus Wunderlich, 1994b: 168 (T from Lepthyphantes)
Lepthyphantes nebulosus Roberts, 1995: 357, f. (mf)
Lepthyphantes nebulosus Zhao, 1995: 871, f. 411a-b (f)
Lepthyphantes nebulosus Roberts, 1998: 369, f. (mf)
Megalepthyphantes nebulosus Paquin & Dupérré, 2003: 143, f. 1577-1579 (mf)
Megalepthyphantes nebulosus Tanasevitch & Saaristo, 2006: 15, f. 5 (f)
Megalepthyphantes nebulosus Saaristo in Marusik & Koponen, 2008a: 8, pl. 1 (m)
Megalepthyphantes nebulosus Dupérré, 2013: 7, f. 63 (m)
Megalepthyphantes nebulosus Danişman & Cosar, 2013: 569, f. 3A-C (f)
Megalepthyphantes nebulosus Marusik, Mikhailov & Omelko, 2015: 119 (S of Lepthyphantes melanopleuros; rejected by Breitling, 2015b: 479, see Linyphia melanopleuros, nomen dubium)
Megalepthyphantes nebulosus Najim & Al-Esa, 2025: 4172, f. 7A-E, 8A-D (f)
Megalepthyphantes pseudocollinus Saaristo, 1997 | | Europe, Russia (Europe to West Siberia), Caucasus, Turkey, Iran urn:lsid:nmbe.ch:spidersp:011588
Lepthyphantes collinus Miller, 1971: 234, pl. 55, f. 20-21 (mf; misidentified per Saaristo, 1997a: 258)
Lepthyphantes collinus Palmgren, 1975: 54, f. 11.14-16 (mf; misidentified per Saaristo, 1997a: 258)
Megalepthyphantes pseudocollinus Saaristo, 1997a: 258, f. 1-4 (Dmf) Original description
Megalepthyphantes pseudocollinus Tanasevitch, 2009a: 401, f. 53-56 (m)
Megalepthyphantes pseudocollinus Türkeş et al., 2015: 563, f. 5A-B (f)
Megalepthyphantes cf. pseudocollinus Otto & Japoshvili, 2018: 381, f. 27 (m)
Megalepthyphantes salam Lecigne, 2025 | | Morocco urn:lsid:nmbe.ch:spidersp:064707
Megalepthyphantes salam Lecigne in Lecigne, Moutaouakil & Lips, 2025: 89, f. 51A-K, 52A-F (Dmf) Original description
Megalepthyphantes sanchaheensis Irfan, Zhang & Peng, 2022 | | China urn:lsid:nmbe.ch:spidersp:057084
Megalepthyphantes sanchaheensis Irfan, Zhang & Peng, 2022: 162, f. 187A-D, 188A-B (Df) Original description
Megalepthyphantes turkestanicus (Tanasevitch, 1989) | | Turkmenistan, Afghanistan, China urn:lsid:nmbe.ch:spidersp:011589
Lepthyphantes turkestanicus Tanasevitch, 1989a: 113, f. 47-51 (Dmf) Original description
Lepthyphantes turkestanicus Song, Zhu & Chen, 1999: 186, f. 103K-L, R (mf)
Megalepthyphantes turkestanicus Saaristo & Tanasevitch, 2000: 264 (T from Lepthyphantes)
Megalepthyphantes turkeyensis Tanasevitch, Kunt & Seyyar, 2005 | | Greece, Cyprus, Turkey urn:lsid:nmbe.ch:spidersp:039916
Lepthyphantes congener Kulczyński, 1908c: 68, pl. 2, f. 17 (m; misidentified per Tanasevitch, 2011b: 74)
Megalepthyphantes turkeyensis Tanasevitch, Kunt & Seyyar, 2005: 279, f. 1-9 (Dm) Original description
Megalepthyphantes turkeyensis Tanasevitch, 2011b: 74, f. 82-83 (m)
Megalepthyphantes turkeyensis Bosmans et al., 2019b: 20, f. 11a-f (m, Df)
Megalepthyphantes turkeyensis Seyyar, Türkeş & Demir, 2022d: 25, f. 3A-D (m)

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