Linyphiidae Blackwall, 1859

Family |   ACCEPTED
N.B.: for comments on relationships, see Michalik & Hormiga, 2010: 7; re-synonymization of Pimoidae (Murphy & Roberts, 2015: ix) not sufficiently justified and is not accepted here; Sinopimoidae Li & Wunderlich, 2008 [urn:lsid:nmbe.ch:spiderfam:0112] synonymyized with Linyphiidae by Dimitrov et al., 2017: 245; resurrection of Sinopimoidae by Wunderlich, 2020b: 20 is not accepted here, because Dimitrov et al., 2017 was not taken into account; re-elevation of Sinopimoidae to family level by Eskov & Marusik, 2023: 343 not accepted here (by majority vote of the WSC boards); the current status of Sinopimoidae is also maintained for reasons of stability and we wait for a probable confirmation or rejection of this resurrection; other subfamilies are Erigoninae Emerton, 1882: 7 and Mynogleninae Lehtinen, 1967: 250.
Gen. Ipa Saaristo, 2007 urn:lsid:nmbe.ch:spidergen:03799
N.B.: although Saaristo indicated that the gender is feminine, he treated all the names as masculine and that was presumably his intention.

In synonymy:

Ipa pirini (Drensky, 1921, removed from S of I. keyserlingi) = Ipa terrenus L. Koch, 1879 (Deltshev, 1988a: 52, sub Lepthyphantes quadrimaculatus, contra Wunderlich, 1980e: 321)

Ipa quadrimaculatus (Kulczyński, 1898, T from Lepthyphantes) = Ipa terrenus L. Koch, 1879 (Saaristo, 2007b: 40)

Ipa keyserlingi (Ausserer, 1867) | | Europe, Caucasus urn:lsid:nmbe.ch:spidersp:011198
Linyphia keyserlingi Ausserer, 1867: 160, pl. 7, f. 1-4 (Dmf) Original description
Linyphia guttata Thorell, 1875b: 61 (Df)
Linyphia guttata Thorell, 1875c: 31 (Df)
Bathyphantes keyserlingi Förster & Bertkau, 1883: 273
Microneta keyserlingi Simon, 1884a: 423, f. 199-201 (mf)
Lephthyphantes keyserlingi Chyzer & Kulczyński, 1894: 71, pl. 3, f. 8 (mf)
Lepthyphantes keyserlingi Bösenberg, 1901: 76, pl. 6, f. 81 (mf)
Lepthyphantes keyserlingi Wiehle, 1956: 210, f. 350-351 (mf)
Lepthyphantes keyserlingi Miller, 1971: 226, pl. XLIV, f. 18-19 (mf)
Lepthyphantes keyserlingi Palmgren, 1975: 62, f. 10.17-19 (mf)
Lepthyphantes keyserlingi Miller & Žitňanská, 1976a: 85, pl. IV, f. 5-7 (mf)
Lepthyphantes keyserlingi Wunderlich, 1980e: 321 (S of Lepthyphantes pirini; rejected by Deltshev, 1988a: 52)
Lepthyphantes keyserlingi Tanasevitch, 1990: 28, f. 11.3-6 (mf)
Lepthyphantes keyserlingi Heimer & Nentwig, 1991: 188, f. 507 (mf)
Lepthyphantes keyserlingi Kronestedt, 1993d: 147, f. 1, 3-4 (f)
Ipa keyserlingi Saaristo, 2007b: 39, f. 3-4, 7-8, 11-12, 14-16 (mf, T from Lepthyphantes)
Ipa keyserlingi Oger & Van Keer, 2017: 19, f. 7A-B (m)
Ipa keyserlingi Roušar, 2022a: 17, f. 3 (f)
Ipa keyserlingi Lecigne, 2023: 15, f. 9A-G (f)
Ipa pepticus (Tanasevitch, 1988) | | Kazakhstan, Turkmenistan, Mongolia urn:lsid:nmbe.ch:spidersp:011274
Lepthyphantes pepticus Tanasevitch, 1988b: 193, f. 33-35 (Dm) Original description
Lepthyphantes pepticus Wunderlich, 1995e: 488, f. 37-38 (m)
Ipa pepticus Saaristo, 2007b: 39, f. 28-29 (m, Df)
Ipa spasskyi (Tanasevitch, 1986) | | Turkey, Ukraine, Russia (Europe, West Siberia), Kazakhstan urn:lsid:nmbe.ch:spidersp:011307
Lepthyphantes spasskyi Tanasevitch, 1986: 140, f. 9-16 (Dmf) Original description
Ipa spasskyi Saaristo, 2007b: 40, f. 26-27 (mf)
Ipa terrenus (L. Koch, 1879) | | Spain, France, Italy, Czechia, Slovakia, Eastern Europe, Turkey, Caucasus, Russia (Europe to South Siberia) urn:lsid:nmbe.ch:spidersp:041363
Linyphia terrena L. Koch, 1879c: 31, pl. 1, f. 19 (Df) Original description
Lepthyphantes quadrimaculatus Kulczyński, 1898: 90, pl. 2, f. 63-64 (Dm) [urn:lsid:nmbe.ch:spidersp:011281]
Lepthyphantes pirini Drensky, 1921: 13, 76, pl. 1. f. 1-4 (Df) [urn:lsid:nmbe.ch:spidersp:057032]
Lepthyphantes terrenus Holm, 1973: 95, f. 76-77 (f, T from Linyphia)
Lepthyphantes quadrimaculatus Miller & Žitňanská, 1976a: 85, pl. 4, f. 1-4 (m, Df, removed from S of L. keyserlingi, rejecting Kulczyński, in Lessert, 1910b: 264)
Lepthyphantes terrenus Tanasevitch & Eskov, 1987: 195 (nomen dubium)
Lepthyphantes quadrimaculatus Deltshev, 1988a: 52, f. 1 (f, S of Lepthyphantes pirini, L. pirini removed from S of L. keyserlingi, rejecting Wunderlich, 1980e: 321)
Lepthyphantes quadrimaculatus Tanasevitch, 1990: 28, f. 11.1-2 (m)
Lepthyphantes quadrimaculatus Heimer & Nentwig, 1991: 188, f. 508 (mf)
Lepthyphantes quadrimaculatus Esyunin, Efimik & Polyanin, 1995: 60, f. 30-32 (mf)
Ipa terrenus Saaristo, 2007b: 40, f. 24-25 (mf, T from Lepthyphantes, removed from nomen dubium, S of Lepthyphantes quadrimaculatus)
Ipa terrenus Oger & Van Keer, 2017: 19, f. 6A-B (m)
Ipa terrenus Naumova & Genchev, 2022: 103, f. 11 (f)
Ipa terrenus Roušar, 2022a: 17, f. 3 (f)
Ipa terrenus Seropian et al., 2023: 249, f. 39-40 (mf)

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