World Spider Catalog

Linyphiidae Blackwall, 1859

Family | ACCEPTED

N.B.: for comments on relationships, see Michalik & Hormiga, 2010: 7; re-synonymization of Pimoidae (Murphy & Roberts, 2015: ix) not sufficiently justified and is not accepted here; Sinopimoidae Li & Wunderlich, 2008 [urn:lsid:nmbe.ch:spiderfam:0112] synonymyized with Linyphiidae by Dimitrov et al., 2017: 245; resurrection of Sinopimoidae by Wunderlich, 2020b: 20 is not accepted here, because Dimitrov et al., 2017 was not taken into account; re-elevation of Sinopimoidae to family level by Eskov & Marusik, 2023: 343 not accepted here (by majority vote of the WSC boards); the current status of Sinopimoidae is also maintained for reasons of stability and we wait for a probable confirmation or rejection of this resurrection; Tian et al., 2022 show five subfamilies: Linyphiinae (incl. Mynogleninae Lehtinen, 1967: 250 and Dubiaraneinae Millidge, 1993c: 145), Erigoninae Emerton, 1882: 7, Micronetinae Hull, 1920a: 8, Ipainae Saaristo, 2007b: 34 and Stemonyphantinae Wunderlich, 1986: 120; da Silva-Moreira, Kulkarni & Hormiga, 2025: 60-62 list resp. newly circumscribe six subfamilies: Erigoninae, Lepthyphantinae Simon, 1929: 562, Linyphiinae (syn. nov.: Dubiaraneinae), Mynogleninae, Pocobletinae Silva-Moreira & Hormiga, 2025: 61 (new subfamily) and Stemonyphantinae, state Ipainae and Micronetinae to be polyphyletic, leave several linyphiid clades unranked and also several genera as incertae sedis.

Gen. Asthenargus Simon & Fage, 1922 urn:lsid:nmbe.ch:spidergen:00851

type Gongylidiellum paganum Simon, 1884; gender masculine

N.B.: not a senior synonym of Jacksonella Millidge, 1951 (Merrett, Locket & Millidge, 1985: 397, contra Millidge, 1977: 40); see Neomaso; not included in a subfamily of Linyphiidae but in an unranked linyphiid clade by da Silva-Moreira, Kulkarni & Hormiga, 2025: 62.

Transferred to other genera:

Asthenargus sexoculatus (Paik & Yaginuma, 1969) -- see Jacksonella

Asthenargus simoni (Lessert, 1904) -- see Trichoncus

Asthenargus tirolensis Schenkel, 1939 -- see Wiehlenarius

In synonymy:

Asthenargus baumi Miller, 1938 = Asthenargus helveticus Schenkel, 1936 (Thaler, 1969: 201)

Asthenargus malickyi (Wunderlich, 1969, T from Gongylidiellum) = Asthenargus bracianus Miller, 1938 (Thaler, 1976a: 228)

Asthenargus adygeicus Tanasevitch, Ponomarev & Chumachenko, 2016 | | Russia (Caucasus) urn:lsid:nmbe.ch:spidersp:048954

Asthenargus adygeicus Tanasevitch, Ponomarev & Chumachenko, 2016b: 423, f. 1-9 (Dmf) Original description

Asthenargus bracianus Miller, 1938 | | Central, Eastern Europe urn:lsid:nmbe.ch:spidersp:009509

Asthenargus bracianus Miller, 1938b: 632, f. 2 (Df) Original description

Gongylidiellum malickyi Wunderlich, 1969: 390, f. 35-36 (Df) [urn:lsid:nmbe.ch:spidersp:057538]

Asthenargus bracianus Thaler, 1976a: 228, f. 1-7 (f, S of Gongylidiellum malickyi)

Asthenargus bracianus Thaler, 1991a: 168, f. 4-6, 8-13 (Dm, f)

Asthenargus bracianus van Helsdingen & IJland, 2015: 26, f. 16-18 (f)

Asthenargus brevisetosus Miller, 1970 | | Angola urn:lsid:nmbe.ch:spidersp:009510

Asthenargus (?) brevisetosus Miller, 1970: 97, pl. XI, f. 1-6 (Df) Original description

Asthenargus carpaticus Weiss, 1998 | | Romania urn:lsid:nmbe.ch:spidersp:009511

Asthenargus carpaticus Weiss, 1998b: 455, f. 1 (Df) Original description

Asthenargus caucasicus Tanasevitch, 1987 | | Caucasus (Russia, Georgia, Azerbaijan) urn:lsid:nmbe.ch:spidersp:009512

Asthenargus caucasicus Tanasevitch, 1987a: 338, f. 74-80 (Dmf) Original description

Asthenargus caucasicus Tanasevitch, 1990: 103, f. 21.26, 22.7, 24.32 (mf)

Asthenargus conicus Tanasevitch, 2006 | | China urn:lsid:nmbe.ch:spidersp:039881

Asthenargus conicus Tanasevitch, 2006b: 289, f. 35-38 (Dmf) Original description

Asthenargus desbaratoensis Wunderlich, 2024 | | Portugal urn:lsid:nmbe.ch:spidersp:062541

Asthenargus desbaratoensis Wunderlich, 2024b: 18, f. 33-35 (Df) Original description

Asthenargus edentulus Tanasevitch, 1989 | | Kazakhstan, Kyrgyzstan, China urn:lsid:nmbe.ch:spidersp:009513

Asthenargus edentulus Tanasevitch, 1989a: 127, f. 117-121 (Dmf) Original description

Asthenargus edentulus Gao, Zhu & Sha, 1994: 80, f. 6-10 (mf)

Asthenargus edentulus Song, Zhu & Chen, 1999: 159, f. 86M-P (mf)

Asthenargus expallidus Holm, 1962 | | Cameroon, DR Congo, Kenya, Tanzania urn:lsid:nmbe.ch:spidersp:009514

Asthenargus expallidus Holm, 1962: 76, f. 21a-d, pl. II, f. 14-15 (Dmf) Original description

Asthenargus helveticus Schenkel, 1936 | | Germany, Poland, Czechia, Switzerland, Austria, Italy urn:lsid:nmbe.ch:spidersp:009515

Asthenargus helveticus Schenkel, 1936a: 317, f. 2 (Df) Original description

Asthenargus baumi Miller, 1938a: 63, f. 8-11 (Dmf) [urn:lsid:nmbe.ch:spidersp:057539]

Asthenargus helveticus Schenkel, 1939: 103, f. 3 (Dm)

Asthenargus baumi Miller, 1951: 245, pl. III, f. 7 (m)

Asthenargus helveticus Thaler, 1969: 201, f. 7-11, 13 (mf, probably junior S of Asthenargus longispina, S of Asthenargus baumi)

Asthenargus helveticus Miller, 1971: 278, pl. LIX, f. 6 (m)

Asthenargus helveticus Millidge, 1977: 40, f. 154 (m)

Asthenargus helveticus Thaler, 1978a: 183, f. 7 (m)

Asthenargus helveticus Heimer & Nentwig, 1991: 116, f. 332 (mf)

Asthenargus inermis Simon & Fage, 1922 | | Kenya, Tanzania urn:lsid:nmbe.ch:spidersp:009516

Asthenargus inermis Simon & Fage, 1922: 544, f. 8(2, 3) (Dm) Original description

Asthenargus inermis Fage & Simon, 1936: 329, f. 19 (Df)

Asthenargus linguatulus Miller, 1970 | | Angola urn:lsid:nmbe.ch:spidersp:009517

Asthenargus linguatulus Miller, 1970: 96, pl. X, f. 1-5 (Df) Original description

Asthenargus longispina (Simon, 1915) | | Spain, France urn:lsid:nmbe.ch:spidersp:009518

Gongylidiellum longispina Simon, 1915a: 478 (Dm; specific epithet is a noun) Original description

Asthenargus longispina Simon, 1926: 459, 525

Asthenargus longispinus Denis, 1962d: 282, f. 8-11 (Df)

Asthenargus longispina Thaler, 1969: 201 (probable senior S of Asthenargus helveticus)

Asthenargus major Holm, 1962 | | Kenya urn:lsid:nmbe.ch:spidersp:009519

Asthenargus major Holm, 1962: 81, f. 23, pl. II, f. 18 (Df) Original description

Asthenargus marginatus Holm, 1962 | | Uganda urn:lsid:nmbe.ch:spidersp:009520

Asthenargus marginatus Holm, 1962: 79, f. 22a-d, pl. II, f. 16-17 (Dmf) Original description

Asthenargus matsudae Saito & Ono, 2001 | | Japan urn:lsid:nmbe.ch:spidersp:037558

Asthenargus matsudae Saito & Ono, 2001: 50, f. 104-107 (Dmf) Original description

Asthenargus matsudae Ono, Matsuda & Saito, 2009: 294, f. 492-495 (mf)

Asthenargus matsudae Suzuki, 2020d: 64, f. 2-3 (mf)

Asthenargus matsudae Hino, 2024: 55, f. 2E-F (mf)

Asthenargus myrmecophilus Miller, 1970 | | Nigeria, Angola urn:lsid:nmbe.ch:spidersp:009521

Asthenargus myrmecophilus Miller, 1970: 94, pl. IX, f. 1-4 (Dm) Original description

Asthenargus myrmecophilus Locket & Russell-Smith, 1980: 74, f. 74-80 (m, Df)

Asthenargus niphonius Saito & Ono, 2001 | | Korea, Japan urn:lsid:nmbe.ch:spidersp:037559

Asthenargus niphonius Saito & Ono, 2001: 48, f. 100-103 (Dmf) Original description

Asthenargus niphonius Ono, Matsuda & Saito, 2009: 294, f. 496-499 (mf)

Asthenargus niphonius Jang et al., 2021a: 296, f. 3A-F (f)

Asthenargus niphonius Serita, 2021e: 274, f. 1p-q (m)

Asthenargus paganus (Simon, 1884) | | Europe, Russia (Europe to West Siberia) urn:lsid:nmbe.ch:spidersp:009522 TYPE SPECIES

Gongylidiellum paganum Simon, 1884a: 602, f. 417-418 (Dmf) Original description

Gongylidiellum paganum Jackson, 1908: 62, pl. 4, f. 14-15 (f)

Asthenargus paganus Simon & Fage, 1922: 545, f. 8(1, 4) (m)

Asthenargus paganus Simon, 1926: 459, 525, f. 790 (mf)

Notioscopus paganus Hull, 1933: 108

Asthenargus paganus Miller, 1936a: 186, f. 5-6 (mf)

Asthenargus paganum Miller, 1951: 245, pl. III, f. 8-9 (m)

Asthenargus paganus Locket & Millidge, 1953: 301, f. 182D-F (mf)

Asthenargus paganus Tullgren, 1955: 333, f. 37a-b (mf)

Asthenargus paganus Wiehle, 1960a: 585, f. 1089-1094 (mf)

Asthenargus paganus Merrett, 1963b: 405, f. 74A-B (m)

Asthenargus paganus Thaler, 1969: 196, f. 12 (f)

Asthenargus paganus Miller, 1971: 278, pl. LIX, f. 5, pl. LX, f. 27 (mf)

Astenargus paganus Palmgren, 1976: 43, f. 10.19-21 (mf)

Asthenargus paganus Millidge, 1977: 40, f. 155, 166 (m)

Asthenargus paganus Thaler, 1978a: 183, f. 8 (m)

Asthenargus paganus Millidge, 1984b: 244, f. 66-67, 81 (f)

Asthenargus paganus Roberts, 1987: 110, f. 54d, h (mf)

Asthenargus paganus Heimer & Nentwig, 1991: 116, f. 333 (mf)

Asthenargus paganus Hormiga, 2000: 30, f. 3A-I (mf)

Asthenargus perforatus Schenkel, 1929 | | France, Italy, Central Europe urn:lsid:nmbe.ch:spidersp:009523

Asthenargus perforatus Schenkel, 1929a: 18, f. 3 (Df) Original description

Asthenargus perforatus Miller, 1951: 243, pl. I, f. 9-13 (f)

Asthenargus perforatus Thaler, 1969: 204, f. 14 (f)

Asthenargus perforatus Miller, 1971: 298, pl. LIX, f. 7-8 (f)

Asthenargus perforatus Moritz, 1973: 184, f. 9-10 (f)

Asthenargus perforatus Thaler, 1978a: 183, f. 1-6 (Dm)

Asthenargus perforatus Heimer & Nentwig, 1991: 116, f. 331 (mf)

Asthenargus placidus (Simon, 1884) | | France urn:lsid:nmbe.ch:spidersp:009524

Gongylidiellum placidum Simon, 1884a: 603, f. 419 (Dm) Original description

Asthenargus placidus Simon, 1926: 458, 525

Asthenargus placidus Lecigne et al., 2022: 17, f. 4A-D (m)

Asthenargus thaleri Wunderlich, 1983 | | Nepal urn:lsid:nmbe.ch:spidersp:009525

Asthenargus thaleri Wunderlich, 1983a: 236, f. 40-46 (Dmf) Original description