Hahniidae Bertkau, 1878

Family |   ACCEPTED
N.B.: the genera of Cryphoecinae Lehtinen, 1967 (authorship correctly Simon, 1898a: 250, sub Cryphoeceae; also Cryphoecini Kishida, 1955: 6 is older) were transferred from the Agelenidae to the Hahniidae by Lehtinen, 1967, considered a subfamily of Dictynidae by Wunderlich, 2004j: 1385 and lately transferred to Cybaeidae by Wheeler et al., 2017: 606; transfers of Cicurina Menge, 1871 and related genera (Cicurininae sensu Lehtinen, 1967) from the Agelenidae to the Dictynidae (Lehtinen, 1967) have been accepted by Cokendolpher, 2004a: 28; Cicurina, as well as Chorizomma Simon, 1872 and Mastigusa Menge, 1854 (which are not explicitely listed by Wheeler et al., 2017) are transferred from the Dictynidae to the Hahniidae by Wheeler et al., 2017: 607; Mastigusa is transferred to the Cybaeidae by Castellucci et al., 2023: 6; the transfers of Chorizomma, Cicurina and Mastigusa to the Cicurinidae by Murphy & Roberts, 2015: viii were not sufficiently justified and are not accepted here; Ono & Ogata, 2018: 619 mention the authorship of these subfamilial taxa: Cicurininae Kishida, 1955: 4, Tetrilini Kishida, 1955: 5 and Chorizommatini Kishida, 1955: 6; but the oldest found mention of Cicurininae is by F. O. Pickard-Cambridge, 1893: 158 (sub Cicurinae, see also Bonnet, 1956: 1091). Cicurinidae new circumscribed (Bromella, Cicurina and Chorizomma) and elevated to family by Gorneau et al., 2023: 11.
Gen. Iberina Simon, 1881 urn:lsid:nmbe.ch:spidergen:01943
N.B.: Lehtinen, 1967: 240 and Wunderlich, 2004j: 1421 transferred several species from Hahnia to Iberina, those transfers have not been accepted by later authors, but after Ledoux, 2014: 31 and were confirmed by Růžička, 2022: 559.

Transferred to other genera:

Iberina spasskyi (Denis, 1958) -- see Hahnia

In synonymy:

Iberina cacuminata (Bösenberg, 1902) = Iberina montana Blackwall, 1841 (Harm, 1966: 354, sub Hahnia)

Iberina caeca (Georgescu & Sarbu, 1992, T from Hahnia) = Iberina microphthalma Snazell & Duffey, 1980 (Růžička, 2022a: 560)

Iberina muscicola (Simon, 1875) = Iberina candida Simon, 1875 (Harm, 1966: 352, sub Hahnia)

Iberina parva (Kulczyński, 1882) = Iberina montana Blackwall, 1841 (Simon, 1937: 1046, sub Hahnia)

Iberina rupicola (Simon, 1884) = Iberina candida Simon, 1875 (Simon, 1937: 1046, sub Hahnia)

Nomen dubium:

Iberina ljovuschkini Pichka, 1965: 1191, f. 1 (f, Russia/Caucasus) -- Růžička, 2022a: 564

Iberina candida (Simon, 1875) | | Canary Is., North Africa, Europe, Turkey, Israel urn:lsid:nmbe.ch:spidersp:021900
Hahnia candida Simon, 1875a: 143 (Df) Original description
Hahnia muscicola Simon, 1875a: 144, pl. 6, f. 1 (Dmf) [urn:lsid:nmbe.ch:spidersp:054090]
Hahnia candida O. Pickard-Cambridge, 1879d: 71 (Dm)
Hahnia rupicola Simon, 1884h: 120 (Dm) [urn:lsid:nmbe.ch:spidersp:054089]
Hahnia candida Bösenberg, 1902: 235, pl. 21, f. 334 (f)
Hahnia muscicola Dahl, 1912b: 596
Hahnia candida Dahl, 1937: 108, f. 12, 23 (f)
Hahnia muscicola Dahl, 1937: 109, f. 9, 27 (m only, f = Hahnia difficilis per Harm, 1966: 356)
Hahnia candida Simon, 1937: 1028, 1046, f. 1599-1600 (mf, S of Hahnia rupicola)
Hahnia candida Locket & Millidge, 1953: 29, f. 20E-F (mf)
Hahnia candida Harm, 1966: 352, f. 17-22 (mf, S of Hahnia muscicola)
Iberina candida Lehtinen, 1967: 240 (T from Hahnia; not accepted by later authors)
Iberina muscicola Lehtinen, 1967: 455, f. 365, 369 (mf, T from Hahnia; not accepted by later authors)
Hahnia candida Roberts, 1985: 168, f. 74d (mf)
Hahnia candida Heimer & Nentwig, 1991: 370, f. 959 (mf)
Hahnia candida Roberts, 1995: 257, f. (mf)
Hahnia candida Roberts, 1998: 273, f. (mf)
Hahnia candida Levy, 2003: 8, f. 9-15 (mf)
Hahnia candida Marusik et al., 2011: 163, f. 13-14, 17 (f)
Hahnia candida Machač & Zedek, 2011: 4, unnumbered f. (mf)
Iberina candida Ledoux, 2014: 33, f. 5, 8A-C, 9 (mf, T from Hahnia per Lehtinen, 1967: 240)
Hahnia candida Machač & Zedek, 2014: 317, f. 1 (mf)
Iberina candida Růžička, 2022a: 559, f. 3C, 4C, 5C (mf)
Iberina difficilis (Harm, 1966) | | France, Italy, Central Europe, Romania, Bulgaria urn:lsid:nmbe.ch:spidersp:021908
Hahnia muscicola Dahl, 1937: 109, f. 14, 25-26 (f; misidentified per Harm, 1966: 356)
Hahnia difficilis Harm, 1966: 356, f. 29-35 (Dmf) Original description
Iberina difficilis Lehtinen, 1967: 240 (T from Hahnia; not accepted by later authors)
Hahnia difficilis Miller, 1971: 180, pl. XXX, f. 8-10 (mf)
Hahnia difficilis Heimer & Nentwig, 1991: 370, f. 961 (mf)
Hahnia difficilis Wiśniewski, Rozwałka & Wesołowska, 2013: 55, f. 1-6 (mf)
Iberina difficilis Ledoux, 2014: 34, f. 10 (f, T from Hahnia per Lehtinen, 1967: 240)
Iberina difficilis Rivera-Quiroz et al., 2020: 5, f. 3e (f; = Harm, 1966: f. 33)
Iberina difficilis Naumova, Blagoev & Deltshev, 2021: 237, f. 9A-C (m)
Iberina difficilis Růžička, 2022a: 559, f. 3E, 4E, 5E (mf)
Iberina difficilis Růžička, 2022b: 3, f. 1A, C (f)
Iberina harmae Wunderlich, 2023 | | Portugal urn:lsid:nmbe.ch:spidersp:059441
Iberina harmae Wunderlich, 2023: 44, f. 79 (Df) Original description
Iberina mazarredoi Simon, 1881 | | Pyrenees (Spain, France) urn:lsid:nmbe.ch:spidersp:021985
Iberina mazarredoi Simon, 1881b: 127 (Dmf) Original description
Iberina mazarredoi Simon, 1898a: 277, f. 274
Iberina mazarredoi Simon, 1937: 1033, 1047, f. 1614 (mf)
Iberina mazarredoi Ledoux, 2014: 32, f. 6A-D (mf)
Iberina mazarredoi Fernández-Pérez, Castro & Prieto, 2014: 80, 7-9 (mf)
Iberina mazarredoi Růžička, 2022a: 560, f. 3A, 4A, 5A (mf)
Iberina microphthalma (Snazell & Duffey, 1980) | | Britain, Switzerland, Germany, Czechia, Slovakia, Hungary, Romania urn:lsid:nmbe.ch:spidersp:021940
Hahnia microphthalma Snazell & Duffey, 1980: 50, f. 1-6 (Df) Original description
Hahnia microphthalma Roberts, 1985: 168, f. 74h (f)
Hahnia microphthalma Heimer & Nentwig, 1991: 368, f. 957 (f)
Iberina caeca Georgescu & Sarbu, 1992: 139, f. 1-6 (Df) [urn:lsid:nmbe.ch:spidersp:021897]
Hahnia caeca Weiss & Sarbu, 1996: 164, f. 1-4 (Dm, T from Iberina)
Hahnia microphthalma Szita et al., 1998: 342, f. 1-2 (f)
Hahnia microphthalma Hänggi & Stäubli, 2012: 63, f. 3 (f)
Iberina microphthalma Ledoux, 2014: 34, f. 11A-E (f, T from Hahnia)
Hahnia candida Machač & Zedek, 2014: 317, f. 2A-B (f; misidentified per Růžička & Dolanský, 2018: 3)
Hahnia microphthalma Růžička & Dolanský, 2014: 5, f. 3a-b (f)
Hahnia microphthalma Růžička & Dolanský, 2016: 45, f. 3a-b (f)
Iberina microphthalma Mezőfi & Markó, 2018: 3, f. 4a-d (f)
Iberina microphthalma Růžička, 2022a: 560, f. 3B, 4B, 5B (mf, S of Hahnia caeca)
Iberina montana (Blackwall, 1841) | | Europe, Turkey, Georgia urn:lsid:nmbe.ch:spidersp:021942
Agelena montana Blackwall, 1841a: 622 (Df; N.B.: primary junior homonym of Agelena montana C. L. Koch, 1834, but that name is a nomen dubium) Original description
Argus montanus Walckenaer, 1847a: 505
Agelena montana Blackwall, 1861a: 157, pl. 10, f. 100 (f, Dm)
Hahnia montana Thorell, 1873: 435
Hahnia parva Kulczyński, 1882b: 30, pl. 3, f. 18 (Dm) [urn:lsid:nmbe.ch:spidersp:054092]
Hahnia cacuminata Bösenberg, 1902: 237, pl. 21, f. 339 (Df) [urn:lsid:nmbe.ch:spidersp:054091]
Hahnia montana Bösenberg, 1902: 238, pl. 21, f. 340 (mf)
Hahnia cacuminata Dahl, 1937: 108, f. 13, 24 (f)
Hahnia montana Dahl, 1937: 113, f. 17 (f; f. 32 = Hahnia pusilla per Harm, 1966: 358)
Hahnia montana Simon, 1937: 1026, 1046, f. 1597-1598 (mf, S of Hahnia parva)
Hahnia cacuminata Hull, 1951: 95, f. 1-3 (f)
Hahnia montana Locket & Millidge, 1953: 28, f. 20B-D (mf)
Hahnia montana Casemir, 1955: 42, f. 5-6 (mf; m is misidentified per Harm, 1966: 354)
Hahnia montana Harm, 1966: 354, f. 23-28 (mf, S of Hahnia cacuminata)
Iberina montana Lehtinen, 1967: 240 (T from Hahnia; not accepted by later authors)
Hahnia montana Loksa, 1969: 121, f. 82B-C, 83C (mf)
Hahnia montana Miller, 1971: 180, pl. XXX, f. 13-15 (mf)
Hahnia montana Roberts, 1985: 168, f. 74c (mf)
Hahnia montana Bürgis, 1988: 11, f. 1-8 (mf)
Hahnia montana Heimer & Nentwig, 1991: 370, f. 960 (mf)
Hahnia montana Roberts, 1995: 254, f. (mf)
Hahnia montana Roberts, 1998: 270, f. (mf)
Hahnia monata Wunderlich, 2004j: 1421, f. 38 (m, lapsus)
Hahnia montana Almquist, 2005: 275, f. 259a-d (mf)
Hahnia montana Marusik & Kunt, 2009a: 78, f. 1-2, 5, 9-10 (mf)
Hahnia montana Marusik et al., 2011: 164, f. 8-10, 15 (f)
Hahnia montana Wiśniewski, Rozwałka & Wesołowska, 2013: 55, f. 17-22 (m)
Iberina montana Ledoux, 2014: 32, f. 7A-C (mf, T from Hahnia per Lehtinen, 1967: 240)
Iberina montana Růžička, 2022a: 562, f. 3D, 4D, 5D (mf)
Iberina montana Růžička, 2022b: 3, f. 1B, D (f)
Iberina montana Seropian et al., 2023: 245, f. 28-30 (mf)
Iberina montana Hirna & Yanul, 2023: 40, f. 4a-b (f)

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