Cybaeidae Banks, 1892

Family |   ACCEPTED
N.B.: traditionally considered a subfamily of Agelenidae, placed as a subfamily of Dictynidae by Lehtinen, 1967 and elevated to family by Forster, 1970b: 13; after Wheeler et al., 2017: 606 Cybaeidae includes Cryphoecinae Lehtinen, 1967 syn. nov. [authorship correctly Simon, 1898a: 250, sub Cryphoeceae; also Cryphoecini Kishida, 1955: 6 is older] (genera Cryphoeca Thorell, 1870, Calymmaria Chamberlin & Ivie, 1937, Ethobuella Chamberlin & Ivie, 1937 and Willisus Roth, 1981 as well as Cryphoecina Deltshev, 1997, Dirksia Chamberlin & Ivie, 1942, Neocryphoeca Roth, 1970 and Tuberta Simon, 1884, which are not explicitely listed by Wheeler et al., 2017; all transferred from Hahniidae) and the genera Blabomma Chamberlin & Ivie, 1937 and Yorima Chamberlin & Ivie, 1942 (transferred from Dictynidae); furthermore Argyroneta Latreille, 1804 (monotypic in Argyronetinae Thorell, 1869) is transferred to the Dictynidae by Wheeler et al., 2017: 607; Mastigusa Menge, 1854 is transferred from the Hahniidae to the Cybaeidae by Castellucci et al., 2023: 6; the transfers of Blabomma, Cryphoeca and Tuberta to Cicurinidae Murphy & Roberts, 2015 [oldest available name is Cicurininae Kishida, 1955: 4] (Murphy & Roberts, 2015: viii) not sufficiently justified and is not accepted here; Cybaeidae were considered a synonym of Argyronetidae Thorell, 1869 by Grothendieck & Kraus, 1994: 272; Cicurinidae is new circumscribed (Bromella, Cicurina and Chorizomma) and elevated to family by Gorneau et al., 2023: 11; included genera of Cybaeidae are listed by Gorneau et al., 2023: 11; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 515, f. 15-16.
Gen. Calymmaria Chamberlin & Ivie, 1937 urn:lsid:nmbe.ch:spidergen:01935
N.B.: Transferred from Agelenidae to Hahniidae by Lehtinen, 1967: 221 and to Cybaeidae by Wheeler et al., 2017: 606. Considered the sister group of Willisus Roth, 1981 and also closely related to Blabomma Chamberlin & Ivie, 1937, Cybaeina Chamberlin & Ivie, 1942, Cybaeota Chamberlin & Ivie, 1933 and Yorima Chamberlin & Ivie, 1942 by Heiss & Draney, 2004: 520.

In synonymy:

Calymmaria cavicola (Banks, 1896) = Calymmaria persica Hentz, 1847 (Heiss & Draney, 2004: 505)

Calymmaria montavencis (Bishop & Crosby, 1926) = Calymmaria persica Hentz, 1847 (Heiss & Draney, 2004: 505)

Calymmaria quadrata (Exline, 1936) = Calymmaria emertoni Simon, 1897 (Heiss & Draney, 2004: 479)

Nomen dubium:

Calymmaria modestella Roewer, 1952 (in Roth, 1952a: 287, replacement name for Tegenaria modesta Banks, 1898b: 230, pl. 14, f. 21, f, Mexico) -- Heiss & Draney, 2004: 461

Calymmaria alleni Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040949
Calymmaria alleni Heiss & Draney, 2004: 469, f. 8-9 (Df) Original description
Calymmaria aspenola Chamberlin & Ivie, 1942 | | USA urn:lsid:nmbe.ch:spidersp:021857
Calymmaria aspenola Chamberlin & Ivie, 1942a: 23, pl. 3, f. 32-34 (Dmf) Original description
Calymmaria aspenola Heiss & Draney, 2004: 470, f. 10-14 (mf)
Calymmaria bifurcata Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040933
Calymmaria emertoni Chamberlin & Ivie, 1937: 215, pl. 3, f. 20-21 (m; misidentified per Heiss & Draney, 2004: 473)
Calymmaria emertoni Exline, 1938: 21, f. 28 (m; misidentified per Heiss & Draney, 2004: 473)
Calymmaria bifurcata Heiss & Draney, 2004: 473, f. 15-19 (Dmf) Original description
Calymmaria californica (Banks, 1896) | | USA urn:lsid:nmbe.ch:spidersp:021858
Tegenaria californica Banks, 1896b: 90 (Dmf) Original description
Calymmaria californica Chamberlin & Ivie, 1937: 213, pl. 1, f. 4-5 (m)
Calymmaria californica Heiss & Draney, 2004: 475, f. 20-24 (mf)
Calymmaria carmel Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040236
Calymmaria carmel Heiss & Draney, 2004: 478, f. 25-27 (Dm) Original description
Calymmaria emertoni (Simon, 1897) | | USA, Canada urn:lsid:nmbe.ch:spidersp:021860
Tegenaria emertoni Simon, 1897c: 17 (Dm) Original description
Tegenaria emertoni Banks, 1913: 181, pl. 13, f. 39-41 (m, Df)
Tegenaria quadrata Exline, 1936b: 22, pl. 1, f. 4 (Df)
Tegenaria californica Exline, 1936b: 25, f. 2 (mf; misidentified per Heiss & Draney, 2004: 479)
Calymmaria emertoni Chamberlin & Ivie, 1937: 215, pl. 2, f. 10-11 (f, not m = C. bifurcata, T from Tegenaria)
Calymmaria emertoni Exline, 1938: 21, f. 3, 29 (f, not m = C. bifurcata)
Calymmaria quadrata Exline, 1938: 23, pl. 4, f. 27 (f)
Calymmaria emertoni Lehtinen, 1967: 454, f. 343, 345 (mf)
Calymmaria emertoni Heiss & Draney, 2004: 479, f. 28-32 (mf, S of Calymmaria quadrata)
Calymmaria farallon Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040237
Calymmaria farallon Heiss & Draney, 2004: 482, f. 33-37 (Dmf) Original description
Calymmaria gertschi Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040238
Calymmaria gertschi Heiss & Draney, 2004: 484, f. 38-39 (Dm) Original description
Calymmaria humboldt Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040239
Calymmaria humboldt Heiss & Draney, 2004: 485, f. 40-42 (Dm) Original description
Calymmaria iviei Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040240
Calymmaria iviei Heiss & Draney, 2004: 486, f. 43-47 (Dmf) Original description
Calymmaria lora Chamberlin & Ivie, 1942 | | USA urn:lsid:nmbe.ch:spidersp:021861
Calymmaria lora Chamberlin & Ivie, 1942a: 22, pl. 3, f. 30-31 (Df) Original description
Calymmaria lora Heiss & Draney, 2004: 489, f. 48-52 (f, Dm)
Calymmaria minuta Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040241
Calymmaria minuta Heiss & Draney, 2004: 493, f. 53-57 (Dmf) Original description
Calymmaria monicae Chamberlin & Ivie, 1937 | | USA urn:lsid:nmbe.ch:spidersp:021863
Calymmaria monicae Chamberlin & Ivie, 1937: 213, pl. 1, f. 1-3, pl. 2, f. 14-15, pl. 3, f. 16-17 (Dmf) Original description
Calymmaria monicae Roth & Brame, 1972: 24, f. 31 (f)
Calymmaria monicae Roth, 1982: 7-8, f. 31 (f)
Calymmaria monicae Roth, 1985: B1-7, f. 31 (f)
Calymmaria monicae Roth, 1994: 53, f. 31 (f)
Calymmaria monicae Heiss & Draney, 2004: 494, f. 58-62 (mf)
Calymmaria monterey Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040242
Calymmaria monterey Heiss & Draney, 2004: 502, f. 63-64 (Df) Original description
Calymmaria nana (Simon, 1897) | | USA, Canada urn:lsid:nmbe.ch:spidersp:021865
Tegenaria nana Simon, 1897c: 17 (Df) Original description
Tegenaria nana Banks, 1913: 189, pl. 13, f. 47 (f)
Tegenaria nana Exline, 1936b: 23, pl. 1, f. 1 (f, Dm)
Calymmaria nana Chamberlin & Ivie, 1937: 215, pl. 2, f. 8-9, pl. 3, f. 22-23 (mf)
Calymmaria nana Exline, 1938: 22, pl. 4, f. 32-33 (mf)
Calymmaria nana Heiss & Draney, 2004: 502, f. 65-69 (mf)
Calymmaria orick Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040243
Calymmaria orick Heiss & Draney, 2004: 503, f. 70-72 (Dm) Original description
Calymmaria persica (Hentz, 1847) | | USA urn:lsid:nmbe.ch:spidersp:039422
Tegenaria persica Hentz, 1847: 463, pl. 24, f. 23 (Dm) Original description
Tegenaria cavicola Banks, 1896d: 203 (Df)
Cybaeus montavencis Bishop & Crosby, 1926: 201, pl. 25, f. 51-53 (Dmf)
Calymmaria cavicola Chamberlin & Ivie, 1937: 213
Calymmaria montavencis Chamberlin & Ivie, 1937: 213
Calymmaria persica Roth, 1952a: 287
Calymmaria perisca Roth & Brown, 1986: 17 (lapsus, nomen dubium)
Calymmaria montavencis Bennett, 1992: 6, f. 20 (f)
Calymmaria persica Heiss & Draney, 2004: 505, f. 73-77 (mf, removed from nomen dubium, S of Calymmaria cavicola and C. montavencis)
Calymmaria rosario Heiss & Draney, 2004 | | Mexico urn:lsid:nmbe.ch:spidersp:040244
Calymmaria rosario Heiss & Draney, 2004: 507, f. 78-79 (Df) Original description
Calymmaria rothi Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040245
Calymmaria rothi Heiss & Draney, 2004: 507, f. 80-84 (Dmf) Original description
Calymmaria scotia Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040246
Calymmaria scotia Heiss & Draney, 2004: 508, f. 85-89 (Dmf) Original description
Calymmaria sequoia Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040247
Calymmaria sequoia Heiss & Draney, 2004: 510, f. 90-92 (Dm) Original description
Calymmaria shastae Chamberlin & Ivie, 1937 | | USA urn:lsid:nmbe.ch:spidersp:021867
Calymmaria shastae Chamberlin & Ivie, 1937: 214, pl. 1, f. 6-7, pl. 3, f. 24 (Dmf) Original description
Calymmaria shastae Heiss & Draney, 2004: 510, f. 93-97 (mf)
Calymmaria sierra Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040248
Calymmaria sierra Heiss & Draney, 2004: 511, f. 98-102 (Dmf) Original description
Calymmaria similaria Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040249
Calymmaria similaria Heiss & Draney, 2004: 513, f. 103-104 (Df) Original description
Calymmaria siskiyou Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040250
Calymmaria siskiyou Heiss & Draney, 2004: 513, f. 105-109 (Dmf) Original description
Calymmaria sueni Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040251
Calymmaria sueni Heiss & Draney, 2004: 514, f. 110-114 (Dmf) Original description
Calymmaria suprema Chamberlin & Ivie, 1937 | | USA, Canada urn:lsid:nmbe.ch:spidersp:021868
Calymmaria suprema Chamberlin & Ivie, 1937: 214, pl. 2, f. 12-13, pl. 3, f. 18-19 (Dmf) Original description
Calymmaria suprema Roth & Brame, 1972: 24, f. 32 (m)
Calymmaria suprema Roth, 1982: 7-8, f. 32 (m)
Calymmaria suprema Roth, 1985: B1-7, f. 32 (m)
Calymmaria suprema Roth, 1994: 53, f. 32 (m)
Calymmaria suprema Heiss & Draney, 2004: 514, f. 115-119 (mf)
Calymmaria tecate Heiss & Draney, 2004 | | Mexico urn:lsid:nmbe.ch:spidersp:040252
Calymmaria tecate Heiss & Draney, 2004: 517, f. 120-122 (Dm) Original description
Calymmaria tubera Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040253
Calymmaria tubera Heiss & Draney, 2004: 518, f. 123-125 (Dm) Original description
Calymmaria virginica Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040254
Calymmaria virginica Heiss & Draney, 2004: 518, f. 126-130 (Dmf) Original description
Calymmaria yolandae Heiss & Draney, 2004 | | USA urn:lsid:nmbe.ch:spidersp:040255
Calymmaria yolandae Heiss & Draney, 2004: 519, f. 131-133 (Dm) Original description

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