Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Odontodrassus Jézéquel, 1965 urn:lsid:nmbe.ch:spidergen:02459
N.B.: see Murphy, 2007: 50, f. 368-369.

In synonymy:

Odontodrassus ciusi (Berland, 1924, T from Drassodes) = Odontodrassus aphanes Thorell, 1897 (Platnick, 1981b: 332, sub O. javanus)

Odontodrassus javanus (Kulczyński, 1911) = Odontodrassus aphanes Thorell, 1897 (Deeleman-Reinhold, 2001: 530)

Odontodrassus monodi (Fage, 1929, T from Poecilochroa) = Odontodrassus mundulus O. Pickard-Cambridge, 1872 (Levy, 1999d: 448)

Odontodrassus nugatorius (Karsch, 1881, T from Drassodes) = Odontodrassus mundulus O. Pickard-Cambridge, 1872 (Bosmans & Hervé, 2021: 60)

Odontodrassus pulumipes (Xu, 1991, T from Drassyllus) = Odontodrassus hondoensis Saito, 1939 (Song, Zhu & Zhang, 2004: 190)

Odontodrassus aphanes (Thorell, 1897) | | Myanmar to Philippines, Japan, New Caledonia, Salomon Is. Introduced to Jamaica, South Africa, Seychelles, Pacific Is. urn:lsid:nmbe.ch:spidersp:026854
Drassus aphanes Thorell, 1897a: 218 (Df; N.B.: apparently omitted by Roewer) Original description
Scotophaeus javanus Kulczyński, 1911b: 470, pl. 21, f. 21, 24 (Df) [urn:lsid:nmbe.ch:spidersp:027661]
Drassodes ciusi Berland, 1924a: 192, f. 54-56 (Dm) [urn:lsid:nmbe.ch:spidersp:054725]
Drassodes ciusi Berland, 1929a: 393, f. 3-4 (Df)
Odontodrassus javanus Platnick, 1981b: 332, f. 1-4 (mf, T from Zelotes per Roewer, S of Drassodes ciusi)
Odontodrassus javanus Kamura, 1990: 32, f. 1-2 (f)
Odontodrassus aphanes Deeleman-Reinhold, 2001: 530, f. 915-921 (mf, T from Drassodes, S of Odontodrassus javanus)
Odontodrassus aphanes Saaristo, 2002: 11, f. 25-27 (mf)
Odontodrassus aphanes Kamura, 2009: 490, f. 91 (f)
Odontodrassus aphanes Saaristo, 2010: 74, f. 11.14-16 (mf)
Odontodrassus aphanes Dierkens & Ramage, 2016: 141, f. 15-16 (f)
Odontodrassus aphanes Azevedo, Griswold & Santos, 2018a: 684, f. 7C (f)
Odontodrassus aphanes Dippenaar-Schoeman et al., 2021C: 40, 6 f. (mf)
Odontodrassus aravaensis Levy, 1999 | | Egypt, Israel, Iraq urn:lsid:nmbe.ch:spidersp:027658
Odontodrassus aravaensis Levy, 1999d: 450, f. 41-42, 47-49 (Dm) Original description
Odontodrassus aravaensis El-Hennawy, 2017a: 171, f. 1-3 (m)
Odontodrassus aravaensis Zamani, Al-Yacoub & Najim, 2022: 144, f. 1D, 2B, 3A (m)
Odontodrassus aravaensis El-Hennawy et al., 2024: 176, f. 3-25 (m, Df)
Odontodrassus bicolor Jézéquel, 1965 | | Ivory Coast urn:lsid:nmbe.ch:spidersp:027659
Odontodrassus bicolor Jézéquel, 1965: 297, f. 5-6 (Dmf) Original description
Odontodrassus hondoensis (Saito, 1939) | | Russia (Far East), China, Korea, Japan urn:lsid:nmbe.ch:spidersp:027660
Iheringia hondoensis Saito, 1939: 35, f. 4(6), pl. 1, f. 9 (Dm) Original description
Iheringia hondoensis Saito, 1959: 44, f. 19a-d (m)
Odontodrassus hondoensis Kamura, 1987d: 30, f. 1-8 (m, Df, T from Iheringia = Otiothops)
Drassyllus pulumipes Xu, 1991b: 38, f. 19-21 (Df) [urn:lsid:nmbe.ch:spidersp:054727]
Odontodrassus hondoensis Paik, 1992d: 164, f. 1-7 (f)
Odontodrassus hondoensis Song, Zhu & Chen, 1999: 453, f. 264J, 265H (mf)
Odontodrassus hondoensis Namkung, 2002: 472, f. 37.13a (f)
Odontodrassus hondoensis Namkung, 2003: 475, f. 37.13a (f)
Odontodrassus hondoensis Song, Zhu & Zhang, 2004: 190, f. 113A-E (mf, S of Drassyllus pulumipes)
Odontodrassus hondoensis Jung et al., 2005: 190, f. 29, 70, 93 (f)
Odontodrassus hondoensis Kamura, 2009: 490, f. 88-90 (mf)
Odontodrassus hondoensis Yin et al., 2012: 1194, f. 637a-g (mf)
Odontodrassus hondoensis Sato, 2012: 68, unnumbered f. (f)
Odontodrassus hondoensis Kim & Lee, 2013: 124, f. 86A-D (mf)
Odontodrassus hondoensis Suzuki & Yu, 2022b: 179, f. 8-9 (m)
Odontodrassus mundulus (O. Pickard-Cambridge, 1872) | | Morocco to Egypt, Israel, Jordan, Iraq, Iran urn:lsid:nmbe.ch:spidersp:027662
Drassus mundulus O. Pickard-Cambridge, 1872: 234, pl. 15, f. 11 (Df) Original description
Drassus nugatorius Karsch, 1881a: 12, pl. 1, f. 10 (Df) [urn:lsid:nmbe.ch:spidersp:026960]
Scotophaeus mundulus Caporiacco, 1928b: 85
Poecilochroa monodi Fage, 1929b: 249, f. 2-3 (Dmf) [urn:lsid:nmbe.ch:spidersp:054726]
Scotophaeus mundulus Caporiacco, 1934a: 134, pl. 1, f. 10 (f)
Poecilochroa monodi Denis, 1966e: 105, f. 4a-b (f)
Odontodrassus mundulus Levy, 1999d: 448, f. 43-46 (mf, T from Scotophaeus, S of Poecilochroa monodi)
Odontodrassus mundulus Bosmans & Hervé, 2021: 60 (S of Drassodes nugatorius)
Odontodrassus mundulus Zamani, Al-Yacoub & Najim, 2022: 144, f. 2C, 3B (m)
Odontodrassus mundulus Zamani, Darvishnia & Marusik, 2023: 353, f. 7B-C, 8A-C (mf)
Odontodrassus muralis Deeleman-Reinhold, 2001 | | Thailand, China, Indonesia (Sulawesi, Lombok) urn:lsid:nmbe.ch:spidersp:037347
Odontodrassus muralis Deeleman-Reinhold, 2001: 533, f. 922-923 (Df) Original description
Odontodrassus nigritibialis Jézéquel, 1965 | | Ivory Coast urn:lsid:nmbe.ch:spidersp:027663
Odontodrassus nigritibialis Jézéquel, 1965: 296, f. 3-4 (Dmf) Original description
Odontodrassus yunnanensis (Schenkel, 1963) | | China urn:lsid:nmbe.ch:spidersp:027910
Scotophaeus yunnanensis Schenkel, 1963: 47, f. 23 (Df) Original description
Odontodrassus yunnanensis Song, Zhu & Zhang, 2004: 192, f. 114A-H (Tf from Scotophaeus, Dm)

Sponsors