Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Macarophaeus Wunderlich, 2011 urn:lsid:nmbe.ch:spidergen:04007

Transferred to other genera:

Macarophaeus sabulum Wunderlich, 2011 -- see Poecilochroa

Macarophaeus cultior (Kulczyński, 1899) | | Canary Is., Madeira urn:lsid:nmbe.ch:spidersp:027861
Scotophaeus cultior Kulczyński, 1899: 350, pl. 6, f. 30 (Dj) Original description
Herpyllus cultior Denis, 1962a: 36, f. 23-25 (Dmf; T from Scotophaeus rejected because of false generic synonymy)
Macarophaeus cultior Wunderlich, 2011: 46, f. 68g (f, T from Scotophaeus)
Macarophaeus insignis Wunderlich, 2011 | | Canary Is. urn:lsid:nmbe.ch:spidersp:043969
Macarophaeus insignis Wunderlich, 2011: 48, f. 89-92, 92a-b (Dmf) Original description
Macarophaeus varius (Simon, 1893) | | Canary Is. urn:lsid:nmbe.ch:spidersp:027907
Scotophaeus varius Simon, 1893a: 371 (Df) Original description
Scotophaeus varius Denis, 1941a: 110, f. 2 (f)
Scotophaeus varius Schmidt, 1956e: 142, f. 1-2 (f, Dm)
Scotophaeus varius Schmidt, 1956g: 241, f. 7b (f)
Scotophaeus cultior Murphy, 2007: 46, f. 292-293 (mf; misidentified per Wunderlich, 2011: 47)
Macarophaeus varius Wunderlich, 2011: 47, f. 66-68, 68a-f (mf, T from Scotophaeus)
Macarophaeus varius García & Suárez, 2019: 72, f. 2d (m)

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