Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Encoptarthria Main, 1954 urn:lsid:nmbe.ch:spidergen:02368
N.B.: transferred from the Prodidomidae to the Gnaphosidae by Murphy, 2007: 50, f. 354-355; not included in the Gnaphosidae (Prodidominae) by Rodrigues & Rheims, 2020b: 661.

In synonymy:

Encoptarthria serventyi Main, 1954 = Encoptarthria echemophthalma Simon, 1908 (Zakharov & Ovtcharenko, 2011: 328)

Encoptarthria echemophthalma (Simon, 1908) | | Australia (Western Australia) urn:lsid:nmbe.ch:spidersp:027480
Megamyrmecion echemophthalmum Simon, 1908d: 396, f. 13 (Df) Original description
Encoptarthria serventyi Main, 1954: 44, pl. 4 f. 1-6 (Df) [urn:lsid:nmbe.ch:spidersp:026429]
Encoptarthria echemophthalma Zakharov & Ovtcharenko, 2011: 327, f. 5a-b (Tf from Megamyrmecion, S, f. m)
Encoptarthria grisea (L. Koch, 1873) | | Australia urn:lsid:nmbe.ch:spidersp:027139
Drassus griseus L. Koch, 1873: 391, pl. 30, f. 8 (Df) Original description
Echemus griseus Rainbow, 1912a: 190
Echemus griseus Rainbow, 1915: 779, pl. 67, f. 6 (Dm)
Echemus griseus Forster, 1979: 48, f. 209-211 (mf)
Encoptarthria grisea Zakharov & Ovtcharenko, 2011: 328, f. 6a-b (Tmf from Megamyrmecion)
Encoptarthria penicillata (Simon, 1908) | | Australia (Western Australia) urn:lsid:nmbe.ch:spidersp:027484
Megamyrmecion penicillatum Simon, 1908d: 394, f. 10 (Dmf) Original description
Encoptarthria penicillata Zakharov & Ovtcharenko, 2011: 328, f. 7a-b (Tmf from Megamyrmecion)
Encoptarthria perpusilla (Simon, 1908) | | Australia (Western Australia) urn:lsid:nmbe.ch:spidersp:027485
Megamyrmecion perpusillum Simon, 1908d: 397, f. 14 (Df) Original description
Encoptarthria perpusilla Zakharov & Ovtcharenko, 2011: 328 (Tf from Megamyrmecion)
Encoptarthria vestigator (Simon, 1908) | | Australia (Western Australia) urn:lsid:nmbe.ch:spidersp:027490
Megamyrmecion vestigator Simon, 1908d: 395, f. 11 (Dm) Original description
Encoptarthria vestigator Zakharov & Ovtcharenko, 2011: 328 (Tf from Megamyrmecion)

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