World Spider Catalog

Gnaphosidae Banks, 1892

Family | ACCEPTED

Gen. Cladothela Kishida, 1928 urn:lsid:nmbe.ch:spidergen:02411

type Cladothela boninensis Kishida, 1928; gender feminine

N.B.: this genus was unknown outside of Japan until the publication of Yaginuma, 1986a: 193; evidently placed as the type genus of Cladothelidae by Kishida, 1930 (Lansania 2: 115-123) but correctly considered a gnaphosid by Yaginuma.

In synonymy:

Cladothela potanini (Schenkel, 1963, T from Phaeocedus) = Cladothela joannisi (Schenkel, 1963) (Song, Zhu & Zhang, 2004: 38)

Cladothela x-notata (Bösenberg & Strand, 1906, T from Drassyllus) = Cladothela unciinsignita (Bösenberg & Strand, 1906) (Kamura, 1991: 55)

Cladothela auster Kamura, 1997 | | Japan urn:lsid:nmbe.ch:spidersp:026830

Cladothela auster Kamura, 1997: 78, f. 1-4 (Dm; f f. 3-4 see Cladothela sakishima) Original description

Cladothela auster Kamura, 2009: 489, f. 81-82 (m; f f. 83 see Cladothela sakishima)

Cladothela auster Kamura, 2025: 36, f. 7A-D, 8A-L, 9A-D (m, Df)

Cladothela bicolor Suzuki, 2022 | | Japan (Ryukyu Is.) urn:lsid:nmbe.ch:spidersp:056011

Cladothela bicolor Suzuki, in Suzuki & Tatsuta, 2022: 320, f. 1A-F, 2A-R, 3A-E (Dmf) Original description

Cladothela bistorta Zhang, Song & Zhu, 2002 | | China urn:lsid:nmbe.ch:spidersp:038046

Cladothela bistorta Zhang, Song & Zhu, 2002a: 243, f. 10-16 (Dm) Original description

Cladothela bistorta Song, Zhu & Zhang, 2004: 37, f. 18A-G (m)

Cladothela boninensis Kishida, 1928 | | Japan urn:lsid:nmbe.ch:spidersp:026831 TYPE SPECIES

Cladothela boninensis Kishida, 1928a: 32, f. 2 (Df) Original description

Cladothela boninensis Yaginuma, 1986a: 193, f. 108.3 (mf)

Cladothela boninensis Kamura, 1991: 50, f. 1-3, 8-20 (mf)

Cladothela boninensis Murphy, 2007: 46, f. 272-273 (mf)

Cladothela boninensis Kamura, 2009: 490, f. 84-87 (mf)

Cladothela boninensis Suzuki et al., 2026b: 125, f. 5A-F (mf)

Cladothela hupingensis Yin, 2012 | | China urn:lsid:nmbe.ch:spidersp:046213

Cladothela hupingensis Yin, in Yin et al., 2012: 1157, f. 615a-c (Df) Original description

Cladothela joannisi (Schenkel, 1963) | | China urn:lsid:nmbe.ch:spidersp:028261

Phaeocedus potanini Schenkel, 1963: 46, f. 22 (Df) [urn:lsid:nmbe.ch:spidersp:027722]

Zelotes joannisi Schenkel, 1963: 59, f. 31a-d (Dm) Original description

Phaeocedus potanini Zhu & Shi, 1985: 153, f. 136a-d (f; belongs to the genus Drassyllus per Zhu & Shi, 1985: 155)

Zelotes joannisi Zhu & Shi, 1985: 155, f. 139a-f (mf, m is misplaced)

Cladothela joannisi Zhang, Song & Zhu, 2002a: 241, f. 1-9 (mf, T from Zelotes)

Cladothela joannisi Song, Zhu & Zhang, 2004: 38, f. 19A-I (mf, S of Phaeocedus potanini)

Cladothela ningmingensis Zhang, Yin & Bao, 2004 | | China urn:lsid:nmbe.ch:spidersp:038995

Cladothela ningmingensis Zhang, Yin & Bao, 2004: 83, f. 1-8 (Dm) Original description

Cladothela notus Kamura, 2025 | | Japan (Ryukyu Is.) urn:lsid:nmbe.ch:spidersp:065822

Cladothela auster Suzuki et al., 2023: 52, f. 9D (f; misidentified per Kamura, 2025: 27)

Cladothela notus Kamura, 2025: 27, f. 2A-J, 3A-G (Dmf) Original description

Cladothela notus Baba & Himuro, 2026: 80, f. 34-37 (mf)

Cladothela notus Suzuki et al., 2026a: 98, f. 2L (f)

Cladothela notus Nagai & Seshima, 2026: 113, f. 1R (m)

Cladothela oculinotata (Bösenberg & Strand, 1906) | | China, Korea, Japan urn:lsid:nmbe.ch:spidersp:026832

Drassodes oculinotatus Bösenberg & Strand, 1906: 119, pl. 16, f. 480 (Df) Original description

Drassodes oculinotatum Yaginuma, 1960: 120, f. 100.1 (f)

Drassodes oculinotatum Yaginuma, 1971: 120, f. 100.1 (f)

Drassodes oculinotatus Paik, 1978e: 414, f. 186 (f)

Drassodes oculinotatus Yaginuma, 1986a: 189, f. 105.3 (mf)

Cladothela oculinotata Kamura, 1991: 51, f. 21-32 (mf, T from Drassodes)

Cladothela oculinotata Paik, 1992h: 36, f. 1-15 (mf)

Cladothela oculinotata Namkung, 2002: 461, f. 37.2a-c (mf)

Cladothela oculinotata Namkung, 2003: 464, f. 37.2a-c (mf)

Cladothela oculinotata Song, Zhu & Zhang, 2004: 40, f. 20A-E (m)

Cladothela oculinotata Jung et al., 2005: 167, f. 2-3, 5182, 101-102 (mf)

Cladothela oculinotata Kamura, 2009: 489, f. 76-80 (mf)

Cladothela oculinotata Kim & Lee, 2013: 85, f. 56A-C (mf)

Cladothela oculinotata Kamura, 2025: 28, f. 1A-B, E-H, 6E (m)

Cladothela parva Kamura, 1991 | | China, Korea, Japan urn:lsid:nmbe.ch:spidersp:026833

Cladothela parva Kamura, 1991: 57, f. 37-43 (Dmf) Original description

Cladothela parva Zhang, Song & Zhu, 2002b: 17, f. 1-5 (m)

Cladothela parva Zhang, Yin & Bao, 2004: 83, f. 9-16 (mf)

Cladothela parva Song, Zhu & Zhang, 2004: 42, f. 21A-F (mf)

Cladothela parva Seo, 2008b: 39, f. 1-4 (mf)

Cladothela parva Kamura, 2009: 488, f. 70-72 (mf)

Cladothela parva Yin et al., 2012: 1158, f. 616a-h (mf)

Cladothela parva Suzuki et al., 2023: 52, f. 9E (f)

Cladothela parva Hidaka, 2023a: 131, f. 5I-J (m)

Cladothela parva Suzuki, 2024b: 87, f. 4-7 (m)

Cladothela sakishima Kamura, 2025 | | Japan (Ryukyu Is.) urn:lsid:nmbe.ch:spidersp:065823

Cladothela auster Kamura, 1997: 78, f. 3-4 (f; misidentified per Kamura, 2025: 33)

Cladothela auster Kamura, 2009: 489, f. 83 (f; misidentified per Kamura, 2025: 33)

Cladothela oculinotata Suzuki & Serita, 2021b: 242, f. 3 (m; misidentified per Kamura, 2025: 33)

Cladothela auster Kamura, 2022c: 26, f. 1I-J (f; misidentified per Kamura, 2025: 33)

Cladothela sakishima Kamura, 2025: 33, f. 5A-J, 6A-D (Dmf) Original description

Cladothela tortiembola Paik, 1992 | | Korea urn:lsid:nmbe.ch:spidersp:026834

Cladothela tortiembola Paik, 1992h: 38, f. 16-24 (Dm) Original description

Cladothela tortiembola Jung et al., 2005: 168, f. 4, 103-104 (m)

Cladothela tortiembola Kim & Lee, 2013: 86, f. 57A-C (m)

Cladothela unciinsignita (Bösenberg & Strand, 1906) | | Korea, Japan urn:lsid:nmbe.ch:spidersp:026835

Drassodes unci-insignitus Bösenberg & Strand, 1906: 120, pl. 16, f. 500 (Dm) Original description

Prosthesima x-notata Bösenberg & Strand, 1906: 122, pl. 16, f. 489 (Df) [urn:lsid:nmbe.ch:spidersp:026834]

Drassodes unciinsignitus Yaginuma, 1986a: 190, f. 105.5 (m)

Zelotes x-notatus Yaginuma, 1986a: 192, f. 106.6 (m; Yaginuma's palpal illustrations indicate the species belongs to Drassyllus rather than Zelotes; cf. Kamura, 1987c: 78)

Drassyllus x-notatus Platnick, 1989b: 471 (T from Zelotes)

Cladothela unciinsignita Kamura, 1991: 55, f. 33-36 (mf, T from Drassodes, S of Drassyllus x-notatus)

Cladothela unciinsignita Seo, 2008b: 42, f. 5-8 (mf)

Cladothela unciinsignita Kamura, 2009: 488, f. 73-75 (mf)

Cladothela unmunensis Seo, 2017 | | Korea urn:lsid:nmbe.ch:spidersp:049560

Cladothela unmunensis Seo, 2017c: 191, f. 1A-F (Df) Original description