Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Drassodex Murphy, 2007 urn:lsid:nmbe.ch:spidergen:03792
Drassodex cervinus (Simon, 1914) | | Pyrenees (Spain, France) urn:lsid:nmbe.ch:spidersp:026872
Drassodes cervinus Simon, 1914: 133, 208 (Dmf) Original description
Drassodex cervinus Murphy, 2007: x (Tmf from Drassodes)
Drassodex cervinus Hervé, Roberts & Murphy, 2009: 17, f. 28-33 (mf)
Drassodex drescoi Hervé, Roberts & Murphy, 2009 | | France, Switzerland, Italy urn:lsid:nmbe.ch:spidersp:042542
Drassodex drescoi Hervé, Roberts & Murphy, 2009: 15, f. 18-19, 24-27 (Dmf) Original description
Drassodex drescoi Hänggi & Stäubli, 2012: 64, f. 7 (f)
Drassodex fritillifer (Simon, 1914) | | Pyrenees (Spain, France) urn:lsid:nmbe.ch:spidersp:026895
Drassodes fritillifer Simon, 1914: 131, 207, f. 215 (Dmf) Original description
Drassodex fritillifer Murphy, 2007: x (Tmf from Drassodes)
Drassodex fritillifer Hervé, Roberts & Murphy, 2009: 18, f. 34-38 (mf)
Drassodex fritillifer Breitling, 2020: 343, f. 11B-C (m)
Drassodex granja Hervé, Roberts & Murphy, 2009 | | Spain urn:lsid:nmbe.ch:spidersp:042543
Drassodes hispanus Simon, 1914: 133, 207, f. 219-222 (mf, misidentified)
Drassodes hispanus Grimm, 1985: 112, f. 121, 128-129 (mf, misidentified)
Drassodes hispanus Heimer & Nentwig, 1991: 418, f. 1105 (mf, misidentified)
Drassodex granja Hervé, Roberts & Murphy, 2009: 21, f. 41-44 (Dmf) Original description
Drassodex heeri (Pavesi, 1873) | | Alps (Switzerland, Italy, Germany, Austria) urn:lsid:nmbe.ch:spidersp:026903
Drassus heerii Pavesi, 1873a: 126, f. 4-6 (Dmf) Original description
Drassus hispanus Lebert, 1877: 223 (m, misidentified)
Drassus heeri Bösenberg, 1902: 295, pl. 27, f. 433 (mf)
Drassodes heeri Lessert, 1905a: 186, f. 1-3 (mf)
Drassodes heeri Lessert, 1910b: 50, f. 41-43 (mf)
Drassodes heeri Simon, 1914: 133, 207, f. 219, 222 (mf; not f. 220-221, =D. simoni)
Drassodes heeri Wiehle, 1967a: 13, f. 48-54 (mf)
Drassodes heeri Grimm, 1985: 109, f. 122, 130-131 (mf; not f. 130c, =D. simoni)
Drassodes heeri Heimer & Nentwig, 1991: 418, f. 1104 (mf)
Drassodex heeri Murphy, 2007: x (Tmf from Drassodes)
Drassodex heeri Hervé, Roberts & Murphy, 2009: 12, f. 10-15 (mf)
Drassodex heeri Hänggi & Stäubli, 2012: 65, f. 7 (f)
Drassodex hispanus (L. Koch, 1866) | | Spain urn:lsid:nmbe.ch:spidersp:026907
Drassus hispanus L. Koch, 1866: 90, pl. 4, f. 60 (Df) Original description
Drassodex hispanus Murphy, 2007: 41, f. 234-235 (Tf from Drassodes)
Drassodex hispanus Hervé, Roberts & Murphy, 2009: 20, f. 39-40 (f)
Drassodex hispanus Breitling, 2020: 343, f. 12A (f)
Drassodex hypocrita (Simon, 1878) | | Europe, Turkey urn:lsid:nmbe.ch:spidersp:026909
Drassus hypocrita Simon, 1878a: 120, pl. 15, f. 8 (Dmf) Original description
Drassus hypocrita Bösenberg, 1902: 296, pl. 28, f. 434 (mf)
Drassodes hypocrita Lessert, 1905a: 191, f. 7-9 (mf)
Drassodes hypocrita Simon, 1914: 121, 132, 207, f. 216-218 (mf)
Drassodes hypocrita Reimoser, 1937a: 15, f. 27, 31 (mf)
Drassodes hypocrita Tystshenko, 1971: 95, f. 169 (m)
Drassodes hypocrita Grimm, 1985: 114, f. 120, 126-127 (mf)
Drassodes hypocrita Heimer & Nentwig, 1991: 418, f. 1103 (mf)
Drassodes hypocrita Roberts, 1998: 108, f. (mf)
Drassodex hypocrita Murphy, 2007: 41 (Tmf from Drassodes; not f. 232-233, =D. simoni)
Drassodex hypocrita Hervé, Roberts & Murphy, 2009: 9, f. 2-9 (mf)
Drassodex hypocrita Wunderlich, 2011: 38, f. 36-39 (mf)
Drassodex hypocrita Akpınar et al., 2011: 16374, f. 2a (f)
Drassodes hypocrita Murphy & Roberts, 2015: 5, f. 3B (f)
Drassodex lesserti (Schenkel, 1936) | | France, Switzerland, Germany urn:lsid:nmbe.ch:spidersp:026908
Drassus hispanus Simon, 1878a: 121, pl. 15, f. 7 (mf, misidentified)
Drassodes hispanus Simon, 1893a: 360, f. 317, 320 (m, misidentified)
Drassodes heeri Lessert, 1904: 283, pl. 5, f. 5 (misidentified)
Drassodes hispanus Lessert, 1905a: 189, f. 4-6 (mf, misidentified)
Drassodes hispanus Lessert, 1910b: 49, f. 38-40 (mf, misidentified)
Drassodes hispanus Simon, 1914: 134, 208, f. 223-226 (mf, misidentified)
Drassodes hispanus lesserti Schenkel, 1936a: 315, f. 1 (Dm) Original description
Drassodex lesserti Hervé, Roberts & Murphy, 2009: 24, f. 49-57 (elevated m from subspecies, Df)
Drassodex lesserti Lecigne, 2018b: 67, f. 6K (f)
Drassodex simoni Hervé, Roberts & Murphy, 2009 | | Alps (France, Italy, Switzerland) urn:lsid:nmbe.ch:spidersp:042541
Drassodes heeri Simon, 1914: 133, 207, f. 220-221 (mf, misidentified)
Drassodes heeri Grimm, 1985: 109, f. 130c (f, misidentified)
Drassodex heeri Murphy, 2007: 41, f. 232-233 (mf, misidentified)
Drassodex simoni Hervé, Roberts & Murphy, 2009: 14, f. 1, 16-17, 20-23 (Dmf) Original description
Drassodex tajikistanicus Fomichev & Marusik, 2021 | | Tajikistan urn:lsid:nmbe.ch:spidersp:053156
Drassodex tajikistanicus Fomichev & Marusik, 2021a: 446, f. 1-8 (Dm) Original description
Drassodex validior (Simon, 1914) | | France urn:lsid:nmbe.ch:spidersp:042544
Drassodes validior Simon, 1914: 134, 208, f. 227 (Dmf) Original description
Drassodex validior Hervé, Roberts & Murphy, 2009: 23, f. 45-48 (removed mf from S of D. hispanus)

Sponsors