Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Cryptodrassus Miller, 1943 urn:lsid:nmbe.ch:spidergen:02414
N.B.: see Murphy, 2007: f. 532-535 for figures of two species from Spain.

In synonymy:

Cryptodrassus jabalpurensis (Gajbe, 2005, T from Drassyllus) = Cryptodrassus khajuriai Tikader & Gajbe, 1976 (Sankaran, Caleb & Sebastian, 2020a: 2)

Cryptodrassus pulchellus Miller, 1943 = Cryptodrassus hungaricus Balogh, 1935 (Weiss, Szinetár & Samu, 1998)

Cryptodrassus beijing Lin & Li, 2022 | | China urn:lsid:nmbe.ch:spidersp:054881
Cryptodrassus beijing Lin & Li in Lin et al., 2022c: 212, f. 12A-C, 13A-B, 14A-D (Dmf) Original description
Cryptodrassus creticus Chatzaki, 2002 | | Greece (Crete), Turkey urn:lsid:nmbe.ch:spidersp:038047
Cryptodrassus creticus Chatzaki in Chatzaki, Thaler & Mylonas, 2002b: 624, f. 55-56, 61-62 (Dmf) Original description
Cryptodrassus creticus Seyyar, Topçu & Demir, 2006: 119, f. 4 (f)
Cryptodrassus creticus Murphy, 2007: 65, f. 530-531 (mf, misplaced in this genus)
Cryptodrassus creticus Çoşkun, Seyyar & Demir, 2022: 132, f. 2A-C, 3A-D (m)
Cryptodrassus helvoloides (Levy, 1998) | | Israel urn:lsid:nmbe.ch:spidersp:028235
Zelotes helvoloides Levy, 1998c: 150, f. 123-125 (Dm), f. 126-127 (f) belongs to C. helvolus per Chatzaki & Russell-Smith, 2017: 240 Original description
Cryptodrassus helvoloides Chatzaki & Russell-Smith, 2017: 240, f. 7-11 (mf, T from Zelotes)
Cryptodrassus helvolus (O. Pickard-Cambridge, 1872) | | Morocco, Algeria, Tunisia, Russia (Europe), Azerbaijan, Turkey, Cyprus, Israel, Iraq, Iran, Kazakhstan urn:lsid:nmbe.ch:spidersp:028236
Melanophora helvola O. Pickard-Cambridge, 1872: 243, pl. 16, f. 23 (Dm) Original description
Prosthesima helvola Simon, 1878a: 98
Zelotes helvolus Levy, 1998c: 148, f. 120-122 (m)
Zelotes helvoloides Levy, 1998c: 150, f. 126-127 (f; misidentified per Chatzaki & Russell-Smith, 2017: 238)
Cryptodrassus helvolus Chatzaki & Russell-Smith, 2017: 238, f. 1-6 (mf, T from Zelotes)
Zelotes helvolus Ponomarev, Prokopenko & Shmatko, 2017: 110, f. 9-10 (m)
Cryptodrassus helvolus Ponomarev et al., 2018a: 246, f. 3-4 (f)
Cryptodrassus helvolus Abdalrazzaq & Najim, 2024: 142, f. 2A-B, 3A-B (f)
Cryptodrassus helvolus Bosmans & Alioua, 2024: 1118, f. 1A-H (mf)
Cryptodrassus helvolus Nuruyeva & Snegovaya, 2024b: 61, f. 1-4 (f)
Cryptodrassus hungaricus (Balogh, 1935) | | France to Greece and Russia (Europe, Caucasus) urn:lsid:nmbe.ch:spidersp:026838
Zelotes hungaricus Balogh, 1935: 18 (Dmf) Original description
Cryptodrassus pulchellus Miller, 1943: 12, pl. I, f. 2-6 (Df) [urn:lsid:nmbe.ch:spidersp:054866]
Cryptodrassus pulchellus Miller, 1947: 55, pl. II, f. 4a-c (f)
Cryptodrassus pulchellus Miller, 1949: 96, f. 13-23 (Dm)
Cryptodrassus pulchellus Miller, 1971: 81, pl. VI, f. 13-14 (m)
Cryptodrassus pulchellus Ovtsharenko, Levy & Platnick, 1994: 3, f. 7-11 (mf)
Cryptodrassus pulchellus Weiss & Moldován, 1998: 522, f. 1-4 (m)
Cryptodrassus hungaricus Weiss, Szinetár & Samu, 1998: 57, f. 1-4 (mf, T from Zelotes, S of Cryptodrassus pulchellus)
Cryptodrassus hungaricus Ledoux & Rollard, 2004: 16, f. 1A-F (mf)
Cryptodrassus hungaricus Trotta, 2005: 164, f. 270-273 (mf)
Cryptodrassus hungaricus Murphy, 2007: 65, f. 528-529 (mf)
Cryptodrassus hungaricus Kovblyuk & Nadolny, 2010: 190, f. 1-19 (mf)
Cryptodrassus hungaricus Wunderlich, 2011: 35, f. 77-80 (mf)
Cryptodrassus hungaricus Naumova, Blagoev & Deltshev, 2021: 233, f. 3A-C (m)
Cryptodrassus hungaricus Ponomarev et al., 2021: 358, f. 16 (f)
Cryptodrassus iranicus Zamani, Chatzaki, Esyunin & Marusik, 2021 | | Iran urn:lsid:nmbe.ch:spidersp:053226
Cryptodrassus iranicus Zamani et al., 2021a: 79, f. 7A-J (Dm) Original description
Cryptodrassus khajuriai (Tikader & Gajbe, 1976) | | India urn:lsid:nmbe.ch:spidersp:027062
Drassyllus khajuriai Tikader & Gajbe, 1976a: 432, f. 5-8 (Df) Original description
Drassyllus khajuriai Tikader, 1982b: 512, f. 490-494 (f)
Drassyllus jabalpurensis Gajbe, 2005d: 136, f. 23-27 (Df) [urn:lsid:nmbe.ch:spidersp:040603]
Drassyllus khajuriai Gajbe, 2007: 489, f. 208-212 (f)
Cryptodrassus khajuriai Sankaran, Caleb & Sebastian, 2020a: 2, f. 1A-D, 2A-D (f, T from Drassyllus, S of Drassyllus jabalpurensis)
Cryptodrassus liyanicus Zamani & Marusik, 2024 | | Iran urn:lsid:nmbe.ch:spidersp:059956
Cryptodrassus liyanicus Zamani & Marusik in Zamani et al., 2024: 295, f. 6A-C (Df) Original description
Cryptodrassus mahabalei (Tikader, 1982) | | India urn:lsid:nmbe.ch:spidersp:027067
Drassyllus mahabalei Tikader, 1982b: 510, f. 485-459 (Df) Original description
Drassyllus mahabalei Gajbe, 2007: 490, f. 213-217 (f)
Cryptodrassus mahabalei Sankaran, Caleb & Sebastian, 2020a: 5, f. 3A-D (f, T from Drassyllus)
Cryptodrassus platnicki (Gajbe, 1987) | | India urn:lsid:nmbe.ch:spidersp:027081
Drassyllus platnicki Gajbe, 1987d: 289, f. 1-5 (Df) Original description
Cryptodrassus platnicki Sankaran, Caleb & Sebastian, 2020a: 7 (T from Drassyllus)
Cryptodrassus ratnagiriensis (Tikader & Gajbe, 1976) | | India urn:lsid:nmbe.ch:spidersp:027088
Drassyllus ratnagiriensis Tikader & Gajbe, 1976a: 431, f. 1-4 (Df) Original description
Drassyllus ratnagiriensis Tikader, 1982b: 514, f. 495-499 (f)
Cryptodrassus ratnagiriensis Sankaran, Caleb & Sebastian, 2020a: 8, f. 4A-D (f, T from Drassyllus)

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