Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Asemesthes Simon, 1887 urn:lsid:nmbe.ch:spidergen:02399
Asemesthes affinis Lessert, 1933 | | Angola urn:lsid:nmbe.ch:spidersp:026647
Asemesthes affinis Lessert, 1933: 99, f. 14 (Df) Original description
Asemesthes albovittatus Purcell, 1908 | | Namibia, South Africa urn:lsid:nmbe.ch:spidersp:026648
Asemesthes albovittatus Purcell, 1908: 245, pl. 11, f. 35 (Df) Original description
Asemesthes albovittatus Dalmas, 1921: 322, f. 123 (f)
Asemesthes albovittatus Tucker, 1923: 286, pl. 8, f. 20 (f, Dm)
Asemesthes albovittatus Dippenaar-Schoeman et al., 2021B: 22, 4 f. (mf)
Asemesthes ales Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:026649
Asemesthes ales Tucker, 1923: 287, pl. 8, f. 21 (Df) Original description
Asemesthes ales Dippenaar-Schoeman et al., 2021B: 23, 1 f. (f)
Asemesthes alternatus Lawrence, 1928 | | Namibia urn:lsid:nmbe.ch:spidersp:026650
Asemesthes alternatus Lawrence, 1928: 228, pl. 21, f. 8 (Df) Original description
Asemesthes ceresicola Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:026651
Asemesthes ceresicola Tucker, 1923: 288, pl. 8, f. 22 (Dmf) Original description
Asemesthes ceresicola Dippenaar-Schoeman et al., 2021B: 24, 5 f. (mf)
Asemesthes decoratus Purcell, 1908 | | Namibia, South Africa urn:lsid:nmbe.ch:spidersp:026652
Asemesthes decoratus Purcell, 1908: 243, pl. 11, f. 30 (Df) Original description
Asemesthes decoratus Dalmas, 1921: 320, f. 125 (f)
Asemesthes decoratus Dippenaar-Schoeman et al., 2021B: 25, 4 f. (f)
Asemesthes flavipes Purcell, 1908 | | Namibia, South Africa urn:lsid:nmbe.ch:spidersp:026653
Asemesthes flavipes Purcell, 1908: 244, pl. 11, f. 32 (Df) Original description
Asemesthes flavipes Dalmas, 1921: 321, f. 127 (f)
Asemesthes flavipes Dippenaar-Schoeman et al., 2021B: 26, 4 f. (f)
Asemesthes fodina Tucker, 1923 | | Namibia, South Africa urn:lsid:nmbe.ch:spidersp:026654
Asemesthes fodina Tucker, 1923: 291, pl. 8, f. 23 (Dm) Original description
Asemesthes fodina Dippenaar-Schoeman et al., 2021B: 27, 3 f. (m)
Asemesthes hertigi Lessert, 1933 | | Angola urn:lsid:nmbe.ch:spidersp:026655
Asemesthes hertigi Lessert, 1933: 98, f. 13 (Df) Original description
Asemesthes kunenensis Lawrence, 1927 | | Namibia urn:lsid:nmbe.ch:spidersp:026656
Asemesthes kunenensis Lawrence, 1927: 12, pl. 1, f. 7 (Df) Original description
Asemesthes lamberti Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:026657
Asemesthes lamberti Tucker, 1923: 292, pl. 8, f. 25 (Df) Original description
Asemesthes lamberti Dippenaar-Schoeman et al., 2021B: 28, 1 f. (f)
Asemesthes lineatus Purcell, 1908 | | Zambia, Namibia, Botswana, South Africa urn:lsid:nmbe.ch:spidersp:026658
Asemesthes lineatus Purcell, 1908: 244, pl. 11, f. 33-34 (Dmf) Original description
Asemesthes lineatus Dalmas, 1921: 319, f. 124, 130 (mf)
Asemesthes lineatus Lawrence, 1927: 11, pl. 1, f. 6 (f)
Asemesthes lineatus Murphy, 2007: 32, f. 100-101 (mf)
Asemesthes lineatus Dippenaar-Schoeman et al., 2021B: 29, 7 f. (mf)
Asemesthes modestus Dalmas, 1921 | | South Africa urn:lsid:nmbe.ch:spidersp:026659
Asemesthes modestus Dalmas, 1921: 318, f. 129 (Dm) Original description
Asemesthes modestus Dippenaar-Schoeman et al., 2021B: 30, 1 f. (m)
Asemesthes montanus Tucker, 1923 | | Namibia, South Africa urn:lsid:nmbe.ch:spidersp:026660
Asemesthes montanus Tucker, 1923: 294, pl. 8, f. 26 (Dmf) Original description
Asemesthes montanus Dippenaar-Schoeman et al., 2021B: 31, p. 31: 6 f., p. 32: 2 f. (mf)
Asemesthes nigristernus Dalmas, 1921 | | South Africa urn:lsid:nmbe.ch:spidersp:026661
Asemesthes nigristernum Dalmas, 1921: 319, f. 123 (Df) Original description
Asemesthes nigristernus Dippenaar-Schoeman et al., 2021B: 33, 1 f. (f)
Asemesthes numisma Tucker, 1923 | | Namibia, South Africa urn:lsid:nmbe.ch:spidersp:026662
Asemesthes numisma Tucker, 1923: 295, pl. 8, f. 27 (Dmf) Original description
Asemesthes numisma Dippenaar-Schoeman et al., 2021B: 34, 5 f. (mf)
Asemesthes oconnori Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:026663
Asemesthes o'connori Tucker, 1923: 297, pl. 8, f. 28 (Dmf) Original description
Asemesthes oconnori Dippenaar-Schoeman et al., 2021B: 35, 3 f. (m)
Asemesthes pallidus Purcell, 1908 | | South Africa urn:lsid:nmbe.ch:spidersp:026664
Asemesthes pallidus Purcell, 1908: 243, pl. 11, f. 31 (Df) Original description
Asemesthes pallidus Dalmas, 1921: 321, f. 126 (f)
Asemesthes pallidus Dippenaar-Schoeman et al., 2021B: 36, 4 f. (f)
Asemesthes paynteri Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:026665
Asemesthes paynteri Tucker, 1923: 298, pl. 8, f. 29 (Df) Original description
Asemesthes paynteri Dippenaar-Schoeman et al., 2021B: 37, 5 f. (f)
Asemesthes perdignus Dalmas, 1921 | | Namibia urn:lsid:nmbe.ch:spidersp:026666
Asemesthes perdignus Dalmas, 1921: 317, f. 122 (Df) Original description
Asemesthes purcelli Tucker, 1923 | | Namibia, South Africa urn:lsid:nmbe.ch:spidersp:026667
Asemesthes purcelli Tucker, 1923: 298, pl. 9, f. 30 (Df) Original description
Asemesthes purcelli Dippenaar-Schoeman et al., 2021B: 38, 5 f. (f)
Asemesthes reflexus Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:026668
Asemesthes reflexus Tucker, 1923: 299, pl. 10, f. 31 (Dmf) Original description
Asemesthes reflexus Dippenaar-Schoeman et al., 2021B: 39, 5 f. (mf)
Asemesthes septentrionalis Caporiacco, 1940 | | Ethiopia urn:lsid:nmbe.ch:spidersp:026669
Asemesthes septentrionalis Caporiacco, 1940c: 836 (Df) Original description
Asemesthes sinister Lawrence, 1927 | | Namibia urn:lsid:nmbe.ch:spidersp:026670
Asemesthes sinister Lawrence, 1927: 13, pl. 1, f. 8 (Df) Original description
Asemesthes subnubilus Simon, 1887 | | Namibia, South Africa urn:lsid:nmbe.ch:spidersp:026671
Asemesthes subnubilus Simon, 1887b: 373 (Dj) Original description
Asemesthes subnubilus Simon, 1893a: 384, f. 340
Asemesthes aureus Purcell, 1908: 242 (Dmf)
Asemesthes subnubilus Dalmas, 1921: 317, f. 4 (j)
Asemesthes aureus Tucker, 1923: 288
Asemesthes subnubilus Tucker, 1923: 301
Asemesthes subnubilus Dippenaar-Schoeman et al., 2021B: 40, 5 f. (m)
Asemesthes subnubilus Haddad et al., 2022: 20, f. 1-13 (mf)
Asemesthes windhukensis Tucker, 1923 | | Namibia urn:lsid:nmbe.ch:spidersp:026672
Asemesthes windhukensis Tucker, 1923: 301, pl. 9, f. 32 (Df) Original description

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