Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Apopyllus Platnick & Shadab, 1984 urn:lsid:nmbe.ch:spidergen:02397

Transferred to other genera:

Apopyllus isabelae Brescovit & Lise, 1993 -- see Nopyllus

In synonymy:

Apopyllus argentina (Mello-Leitão, 1944, T from Eilica) = Apopyllus silvestrii Simon, 1905 (Platnick & Shadab, 1984: 3)

Apopyllus iheringi (Mello-Leitão, 1943) = Apopyllus suavis Simon, 1893 (Azevedo et al., 2016: 306)

Apopyllus melanophorus (Mello-Leitão, 1941, T from Zelotes) = Apopyllus silvestrii Simon, 1905 (Platnick & Shadab, 1984: 4)

Apopyllus pauper (Mello-Leitão, 1942) = Apopyllus suavis Simon, 1893 (Azevedo et al., 2016: 306)

Apopyllus unanimus (Mello-Leitão, 1947, T from Zelotes) = Apopyllus suavis Simon, 1893 (Platnick, 1993c: 642)

Apopyllus aeolicus Azevedo, Ott, Griswold & Santos, 2016 | | Brazil urn:lsid:nmbe.ch:spidersp:048804
Apopyllus aeolicus Azevedo et al., 2016: 312, f. 3E, 4E, 5E, 6E, 7E, 8E, 16F (Dm) Original description
Apopyllus atlanticus Azevedo, Ott, Griswold & Santos, 2016 | | Brazil urn:lsid:nmbe.ch:spidersp:048806
Apopyllus atlanticus Azevedo et al., 2016: 320, f. 3G, 4G, 5G, 6G, 7G, 8G, 9E-F, 10H, 11H, 12H, 13H (Dmf) Original description
Apopyllus centralis Azevedo, Ott, Griswold & Santos, 2016 | | Brazil, Paraguay urn:lsid:nmbe.ch:spidersp:048805
Apopyllus centralis Azevedo et al., 2016: 314, f. 3F, 4F, 5F, 6F, 7F, 8F, 10G, 11G, 12G, 13G, 16D (Dmf) Original description
Apopyllus centralis Espejo, Kochalka & Barrett, 2019: 598, f. 4a-e, 5a-e (f)
Apopyllus gandarela Azevedo, Ott, Griswold & Santos, 2016 | | Brazil urn:lsid:nmbe.ch:spidersp:048803
Apopyllus gandarela Azevedo et al., 2016: 311, f. 10F, 11F, 12F, 13F, 16E (Df) Original description
Apopyllus gandarela Rivera-Quiroz et al., 2020: 5, f. 3d (f; = Azevedo et al., 2016: f. 10F)
Apopyllus huanuco Platnick & Shadab, 1984 | | Peru urn:lsid:nmbe.ch:spidersp:026637
Apopyllus huanuco Platnick & Shadab, 1984: 6, f. 9-10 (Df) Original description
Apopyllus huanuco Azevedo et al., 2016: 309, f. 10D, 11D, 12D, 13D (f)
Apopyllus ivieorum Platnick & Shadab, 1984 | | Mexico urn:lsid:nmbe.ch:spidersp:026640
Apopyllus ivieorum Platnick & Shadab, 1984: 8, f. 15-16 (Dm) Original description
Apopyllus ivieorum Azevedo et al., 2016: 310, f. 3D, 4D, 5D, 6D, 7D, 8D (m)
Apopyllus kanguery Espejo, Kochalka & Barrett, 2019 | | Paraguay urn:lsid:nmbe.ch:spidersp:051395
Apopyllus kanguery Espejo, Kochalka & Barrett, 2019: 596, f. 1a-d, 2 (Df) Original description
Apopyllus malleco Platnick & Shadab, 1984 | | Chile urn:lsid:nmbe.ch:spidersp:026641
Apopyllus malleco Platnick & Shadab, 1984: 6, f. 7-8 (Df) Original description
Apopyllus malleco Azevedo et al., 2016: 309, f. 1A-B, 10C, 11C, 12C, 13C (mf)
Apopyllus now Platnick & Shadab, 1984 | | Curaçao, Colombia urn:lsid:nmbe.ch:spidersp:026642
Apopyllus now Platnick & Shadab, 1984: 7, f. 19-22 (Dmf) Original description
Apopyllus now Azevedo et al., 2016: 310, f. 3C, 4C, 5C, 6C, 7C, 8C, 10E, 11E, 12E, 13E (mf)
Apopyllus silvestrii (Simon, 1905) | | Peru, Bolivia, Brazil, Paraguay, Argentina, Chile urn:lsid:nmbe.ch:spidersp:026644
Melanophora silvestrii Simon, 1905e: 4 (Df) Original description
Zelotes silvestrii Petrunkevitch, 1911: 151
Zelotes melanophorus Mello-Leitão, 1941d: 169, f. 59 (Dmf) [urn:lsid:nmbe.ch:spidersp:060754]
Gytha argentina Mello-Leitão, 1944b: 351, f. 42 (Dm) [urn:lsid:nmbe.ch:spidersp:060753]
Apopyllus silvestrii Platnick & Shadab, 1984: 3, f. 1-4 (mf, T from Zelotes, S of Eilica argentina and Zelotes melanophorus)
Apopyllus silvestrii Murphy, 2007: 50, f. 340-341 (mf)
Apopyllus silvestrii Ott, 2014b: 259, f. 41-47 (m)
Apopyllus silvestrii Azevedo et al., 2016: 305, f. 2A-D, 3A, 4A, 5A, 6A, 7A, 8A, 9A-B, 10A, 11A, 12A, 13A (mf)
Apopyllus silvestrii Azevedo, Griswold & Santos, 2018a: 679, f. 2C, 3D (m)
Apopyllus silvestrii Espejo, Kochalka & Barrett, 2019: 597, f. 3a-f (m)
Apopyllus suavis (Simon, 1893) | | Venezuela, Peru, Brazil, Argentina urn:lsid:nmbe.ch:spidersp:026645
Herpyllus suavis Simon, 1893b: 455, pl. 9, f. 25 (Df; lectotype only, paralectotype belongs to A. centralis, per Azevedo et al., 2016: 314) Original description
Zelotes pauper Mello-Leitão, 1942b: 413, f. 39 (Df) [urn:lsid:nmbe.ch:spidersp:026643]
Zelotes iheringi Mello-Leitão, 1943b: 262 (Df) [urn:lsid:nmbe.ch:spidersp:026638]
Zelotes unanimus Mello-Leitão, 1947b: 280, pl. 38, f. 37 (Df) [urn:lsid:nmbe.ch:spidersp:060755]
Herpyllus cf. suavis Brignoli, 1972c: 380, f. 38, 41 (f)
Apopyllus iheringi Platnick & Shadab, 1984: 5, f. 11-14 (Dm, f, T from Zelotes)
Apopyllus pauper Platnick & Shadab, 1984: 4, f. 5-6 (f, T from Zelotes)
Apopyllus suavis Platnick & Shadab, 1984: 7, f. 17-18 (f, T from Herpyllus)
Apopyllus iheringi Platnick, 1993c: 642 (S of Zelotes unanimus)
Apopyllus suavis Azevedo et al., 2016: 306, f. 3B, 4B, 5B, 6B, 7B, 8B, 9C-D, 10B, 11B, 12B, 13B, 16A-C (mf, S of Apopyllus pauper and A. iheringi)

Sponsors