World Spider Catalog

Entypesidae Bond, Opatova & Hedin, 2020

Family | ACCEPTED

N.B.: new family by Bond, Opatova & Hedin, in Opatova et al., 2020: 702, including three genera formerly in Nemesiidae (Entypesa, Hermacha and Lepthercus).

In synonymy:

Damarchodes Simon, 1903 = Hermacha Simon, 1889 (Tucker, 1917: 112, confirmed by Raven, 1985a: 85)

Pseudohermacha Strand, 1907 = Entypesa Simon, 1902 (Raven, 1985a: 86)

Gen. Afropesa Zonstein & Ríos-Tamayo, 2021 urn:lsid:nmbe.ch:spidergen:04720

Afropesa gauteng Zonstein & Ríos-Tamayo, 2021 | | South Africa urn:lsid:nmbe.ch:spidersp:053106

Afropesa gauteng Zonstein & Ríos-Tamayo, 2021: 17, f. 19-36 (Dmf) Original description

Afropesa schoutedeni (Benoit, 1965) | | South Africa urn:lsid:nmbe.ch:spidersp:001121

Entypesa schoutedeni Benoit, 1965b: 261, f. 3-7 (Dmf) Original description

Entypesa schoutedeni Raven, 1983b: 552, pl. I, f. 1-2, pl. II, f. 1

Entypesa schoutedeni Raven, 1985a: 86, f. 41 (m)

Afropesa schoutedeni Zonstein & Ríos-Tamayo, 2021: 13, f. 1-18 (mf, T from Entypesa)

Afropesa schwendingeri Zonstein & Ríos-Tamayo, 2021 | | South Africa urn:lsid:nmbe.ch:spidersp:053107

Afropesa schwendingeri Zonstein & Ríos-Tamayo, 2021: 23, f. 37-52 (Dmf) Original description

Gen. Brachytheliscus Pocock, 1902 urn:lsid:nmbe.ch:spidergen:04742

N.B.: considered a junior synonym of Hermacha Simon, 1889 by Hewitt, 1915d: 125, revalidated by Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 68.

Brachytheliscus bicolor Pocock, 1897 | | South Africa urn:lsid:nmbe.ch:spidersp:001124

Brachythele bicolor Pocock, 1897b: 735 (Df) Original description

Brachytheliscus bicolor Pocock, 1902b: 317 (Dm)

Hermacha bicolor Hewitt, 1915d: 125

Hermacha bicolor Tucker, 1917: 107, f. 5 (mf)

Brachytheliscus bicolor Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 70, f. 42A-E, 43A,F, 44A-F (mf, T from Hermacha)

Gen. Ekapa Ríos-Tamayo, Lyle & Sole, 2023 urn:lsid:nmbe.ch:spidergen:05929

In synonymy:

Ekapa nigrum (Tucker, 1917, T from Hermacha) = Ekapa curvipes Purcell, 1902 (Ríos-Tamayo, Lyle & Sole, 2023: 4)

Ekapa curvipes (Purcell, 1902) | | South Africa urn:lsid:nmbe.ch:spidersp:001130

Hermacha curvipes Purcell, 1902b: 377 (Dm) Original description

Hermacha curvipes Tucker, 1917: 106 (Df)

Hermacha nigra Tucker, 1917: 110 (Df) [urn:lsid:nmbe.ch:spidersp:043902]

Damarchodes purcelli Bonnet, 1956: 1373 (regarded Hermacha nigra as S, but none of the cited comments of Hewitt, 1919b: 109 justify that conclusion, see also Tucker, 1917: 112)

Hermacha curvipes Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 74, f. 45A-D (m, incertae sedis, differs from currently described genera)

Hermacha nigra Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 75, f. 45E (f, incertae sedis, differs from currently described entypesid genera from South Africa)

Ekapa curvipes Ríos-Tamayo, Lyle & Sole, 2023: 4, f. 1A-G, 2A-E, 3A-F, 4A-I, 5A-E (mf, T from Hermacha, S of Hermacha nigra)

Gen. Entypesa Simon, 1902 urn:lsid:nmbe.ch:spidergen:00114

N.B.: transferred from the Dipluridae to the Nemesiidae and considered a senior synonym of Pseudohermacha Strand, 1907a: 549 [urn:lsid:nmbe.ch:spidergen:04937] (type P. annulipes Strand, 1907) by Raven, 1985a: 86; transferred to the Entypesidae by Opatova et al., 2020: 702.

Transferred to other genera:

Entypesa schoutedeni Benoit, 1965 -- see Afropesa

Nomen dubium:

Entypesa annulipes (Strand, 1907a: 549, Strand, 1907f: 184, j, Madagascar, originally in Pseudohermacha) -- Nentwig et al., 2020: 24

Entypesa andohahela Zonstein, 2018 | | Madagascar urn:lsid:nmbe.ch:spidersp:050627

Entypesa andohahela Zonstein, 2018c: 471, f. 1-16 (Dmf) Original description

Entypesa enakara Zonstein, 2018 | | Madagascar urn:lsid:nmbe.ch:spidersp:050628

Entypesa enakara Zonstein, 2018c: 471, f. 17-27 (Dm) Original description

Entypesa fisheri Zonstein, 2018 | | Madagascar urn:lsid:nmbe.ch:spidersp:050629

Entypesa fisheri Zonstein, 2018c: 474, f. 28-33 (Df) Original description

Entypesa nebulosa Simon, 1902 | j | Madagascar urn:lsid:nmbe.ch:spidersp:001120

Entypesa nebulosa Simon, 1902g: 599 (Df) Original description

Entypesa nebulosa Zonstein, 2018c: 469 (holotype is subadult)

Entypesa rastellata Zonstein, 2018 | | Madagascar urn:lsid:nmbe.ch:spidersp:050630

Entypesa rastellata Zonstein, 2018c: 476, f. 38-50 (Dm) Original description

Gen. Hermacha Simon, 1889 urn:lsid:nmbe.ch:spidergen:00116

N.B.: transferred from the Ctenizidae to the Nemesiidae by Raven, 1985a: 85 and to the Entypesidae by Opatova et al., 2020: 702; considered a senior synonym of Brachytheliscus Pocock, 1902b: 317 by Hewitt, 1915d: 125, of Damarchodes Simon, 1903k: 43 [urn:lsid:nmbe.ch:spidergen:04938] (type D. purcelli Simon, 1903) by Purcell in Tucker, 1917: 112 and of Hermachola Hewitt, 1915 by Raven, 1985a: 85; Brachytheliscus and Hermachola were revalidated by Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 56, 68; the inclusion of the South American species in the genus is uncertain after Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 3.

Transferred to other genera:

Hermacha bicolor (Pocock, 1897) -- see Brachytheliscus

Hermacha capensis (Ausserer, 1871) -- see Hermachola

Hermacha crudeni Hewitt, 1913 -- see Hermachola

Hermacha curvipes Purcell, 1902 -- see Ekapa

Hermacha grahami (Hewitt, 1915) -- see Hermachola

Hermacha iricolor Mello-Leitão, 1923 -- see Rachias

Hermacha leporina Simon, 1891 -- see Stenoterommata

Hermacha nigra Tucker, 1917 -- see Ekapa

In synonymy:

Hermacha capensis (Zonstein, 2016, T from Pionothele) = Hermacha brevicauda Purcell, 1903 (Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 34)

Homonym replaced:

Hermacha purcelli Tucker, 1917 -- see Hermacha tuckeri Raven, 1985

Nomen dubium:

Hermacha nigromarginata Strand, 1907a: 548, Strand, 1907f: 181 (m, South Africa) -- Nentwig et al., 2020: 24

Hermacha anomala (Bertkau, 1880) | | Brazil urn:lsid:nmbe.ch:spidersp:001123

Nemesia anomala Bertkau, 1880a: 17, pl. 1, f. 3 (Df) Original description

Hermacha anomala Simon, 1892a: 114

Hermacha brevicauda Purcell, 1903 | | South Africa urn:lsid:nmbe.ch:spidersp:001125

Hermacha brevicauda Purcell, 1903b: 98 (Dm) Original description

Pionothele capensis Zonstein, 2016a: 36, f. 9-20 (Dmf) [urn:lsid:nmbe.ch:spidersp:048499]

Hermacha brevicauda Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 34, f. 2A, 5A-D, 20A-F, 21A-E (m, S of Pionothele capensis)

Hermacha caudata Simon, 1889 | | Mozambique urn:lsid:nmbe.ch:spidersp:001127

Hermacha caudata Simon, 1889h: 408 (Dm) Original description

Hermacha caudata Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 37, f. 5E-H, 22A-E, 23A-E (m)

Hermacha conspersa Mello-Leitão, 1941 | | Colombia urn:lsid:nmbe.ch:spidersp:001128

Hermacha conspersa Mello-Leitão, 1941b: 234 (Df) Original description

Hermacha evanescens Purcell, 1903 | | South Africa urn:lsid:nmbe.ch:spidersp:001131

Hermacha evanescens Purcell, 1903b: 99 (Df) Original description

Hermacha evanescens Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 39, f. 1B-C, 2B, 3A-D, 5I-L, 24A-F, 25A-E, 26A-E (Dm, f)

Hermacha fossor (Bertkau, 1880) | | Brazil urn:lsid:nmbe.ch:spidersp:001132

Nemesia fossor Bertkau, 1880a: 19, pl. 1, f. 4 (Df) Original description

Hermacha fossor Simon, 1892a: 114

Hermacha fulva Tucker, 1917 | | South Africa urn:lsid:nmbe.ch:spidersp:001133

Hermacha fulva Tucker, 1917: 109, f. 6 (Dm) Original description

Hermacha fulva Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 11, f. 2C, 4A-D, 6A-F, 7A-E (m)

Hermacha itatiayae Mello-Leitão, 1923 | | Brazil urn:lsid:nmbe.ch:spidersp:001136

Hermacha itatiayae Mello-Leitão, 1923a: 339 (Df) Original description

Hermacha lanata Purcell, 1902 | | South Africa urn:lsid:nmbe.ch:spidersp:001137

Hermacha lanata Purcell, 1902b: 373 (Dm) Original description

Hermacha lanata Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 44, f. 2D, 5M-P, 27A-F, 28A-E, 29A-E (m, Df)

Hermacha maraisae Ríos-Tamayo, Engelbrecht & Goloboff, 2021 | | South Africa urn:lsid:nmbe.ch:spidersp:053426

Hermacha maraisae Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 49, f. 2E, 5Q-T, 30A-F, 31A-E (Dm) Original description

Hermacha mazoena Hewitt, 1915 | | Zimbabwe urn:lsid:nmbe.ch:spidersp:001138

Hermacha mazoena Hewitt, 1915c: 312 (Df) Original description

Hermacha montana Ríos-Tamayo, Engelbrecht & Goloboff, 2021 | | South Africa urn:lsid:nmbe.ch:spidersp:053425

Hermacha montana Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 13, f. 2F, 4E-H, 8A-F, 9A-E, 10A-E (Dmf) Original description

Hermacha nigrispinosa Tucker, 1917 | | South Africa urn:lsid:nmbe.ch:spidersp:001139

Hermacha nigrispinosus Tucker, 1917: 112, f. 7 (Dmf) Original description

Hermacha nigrispinosa Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 21, f. 2G, 4I-L, 11A-E, 12A-E, 13A-E (mf)

Hermacha purcelli (Simon, 1903) | j | South Africa urn:lsid:nmbe.ch:spidersp:001141

Damarchodes purcelli Simon, 1903k: 43 (Dj) Original description

Hermacha purcelli Purcell in Tucker, 1917: 112 (T from Damarchodes)

Hermacha septemtrionalis Ríos-Tamayo, Engelbrecht & Goloboff, 2021 | | South Africa urn:lsid:nmbe.ch:spidersp:053429

Hermacha septemtrionalis Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 51, f. 1D-E, 5U-X, 32A-F, 33A-E, 34A-F (Dmf) Original description

Hermacha sericea Purcell, 1902 | | South Africa urn:lsid:nmbe.ch:spidersp:001142

Hermacha sericea Purcell, 1902b: 375 (Dm) Original description

Hermacha sericea Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 24, f. 2H, 4M-P, 14A-F, 15A-J, 16A-E (m, Df)

Hermacha tuckeri Raven, 1985 | | South Africa urn:lsid:nmbe.ch:spidersp:001143

Hermacha purcelli Tucker, 1917: 114, f. 8 (Dmf; preoccupied by Simon, 1903, see Hermacha purcelli) [urn:lsid:nmbe.ch:spidersp:056000]

Hermacha tuckeri Raven, 1985a: 161 (replacement name) Original description

Hermacha tuckeri Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 30, f. 1F, 2I, 4Q-T, 17A-F, 18A-E, 19A-E (mf)

Gen. Hermachola Hewitt, 1915 urn:lsid:nmbe.ch:spidergen:04774

N.B.: considered a junior synonym of Hermacha Simon, 1889 by Raven, 1985a: 85, revalidated by Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 56.

In synonymy:

Hermachola grahami Hewitt, 1915 = Hermachola crudeni Hewitt, 1913 (Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 60)

Hermachola capensis (Ausserer, 1871) | | South Africa urn:lsid:nmbe.ch:spidersp:001126

Brachythele capensis Ausserer, 1871a: 175 (Df) Original description

Hermacha capensis Benoit, 1964e: 417 (Tmf from Brachythele, after Simon, 1903a: 906)

Hermachola capensis Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 58, f. 35A-F (f, T from Hermacha)

Hermachola crudeni (Hewitt, 1913) | | South Africa urn:lsid:nmbe.ch:spidersp:001129

Hermacha crudeni Hewitt, 1913c: 466 (Df) Original description

Hermachola grahami Hewitt, 1915c: 314, f. 7A-C (Dm) [urn:lsid:nmbe.ch:spidersp:001134]

Hermachola crudeni Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 60, f. 3E-H, 36A-E, 27A-D, 38A-F (mf, T from Hermacha, S of Hermacha grahami)

Hermachola lyleae Ríos-Tamayo, Engelbrecht & Goloboff, 2021 | | South Africa urn:lsid:nmbe.ch:spidersp:053428

Hermachola lyleae Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 63, f. 1A, 39A-E, 40A-D, 41A-E (Dmf) Original description

Gen. Lepthercus Purcell, 1902 urn:lsid:nmbe.ch:spidergen:00120

N.B.: transferred from the Ctenizidae to the Nemesiidae by Raven, 1985a: 86 and to the Entypesidae by Opatova et al., 2020: 702.

Lepthercus confusus Ríos-Tamayo & Lyle, 2020 | | South Africa urn:lsid:nmbe.ch:spidersp:051813

Lepthercus confusus Ríos-Tamayo & Lyle, 2020: 269, f. 3A-I, 4A-E (Dmf) Original description

Lepthercus dippenaarae Ríos-Tamayo & Lyle, 2020 | | South Africa urn:lsid:nmbe.ch:spidersp:051818

Lepthercus dippenaarae Ríos-Tamayo & Lyle, 2020: 285, f. 2B, 14A-J, 15A-E (Dmf) Original description

Lepthercus dregei Purcell, 1902 | | South Africa urn:lsid:nmbe.ch:spidersp:001161

Lepthercus dregei Purcell, 1902b: 379 (Dm) Original description

Lepthercus dregei Raven, 1985a: 86, f. 44-49 (m)

Lepthercus dregei Ríos-Tamayo & Lyle, 2020: 272, f. 2J, 5A-J, 6A-D, 7A-C (m, Df)

Lepthercus engelbrechti Ríos-Tamayo & Lyle, 2020 | | South Africa urn:lsid:nmbe.ch:spidersp:051819

Lepthercus engelbrechti Ríos-Tamayo & Lyle, 2020: 288, f. 2H-I, 16A-B, 17A-K, 18A-C (Dmf) Original description

Lepthercus filmeri Ríos-Tamayo & Lyle, 2020 | | South Africa urn:lsid:nmbe.ch:spidersp:051814

Lepthercus filmeri Ríos-Tamayo & Lyle, 2020: 276, f. 2E, 8A-E (Df) Original description

Lepthercus haddadi Ríos-Tamayo & Lyle, 2020 | | South Africa urn:lsid:nmbe.ch:spidersp:051820

Lepthercus haddadi Ríos-Tamayo & Lyle, 2020: 292, f. 2A,F-G, 19A-K, 20A-C (Dmf) Original description

Lepthercus kwazuluensis Ríos-Tamayo & Lyle, 2020 | | South Africa urn:lsid:nmbe.ch:spidersp:051815

Lepthercus kwazuluensis Ríos-Tamayo & Lyle, 2020: 277, f. 2C, 9A-J, 10A-D (Dmf) Original description

Lepthercus lawrencei Ríos-Tamayo & Lyle, 2020 | | South Africa urn:lsid:nmbe.ch:spidersp:051816

Lepthercus lawrencei Ríos-Tamayo & Lyle, 2020: 280, f. 2D, 11A-D (Df) Original description

Lepthercus mandelai Ríos-Tamayo & Lyle, 2020 | | South Africa urn:lsid:nmbe.ch:spidersp:051817

Lepthercus mandelai Ríos-Tamayo & Lyle, 2020: 282, f. 2D, 12A-J, 13A-D (Dmf) Original description

Lepthercus rattrayi Hewitt, 1917 | | South Africa urn:lsid:nmbe.ch:spidersp:001162

Lepthercus rattrayi Hewitt, 1917: 699, pl. 47, f. 1-2, f. 3A-E (Dmf) Original description

Lepthercus rattrayi Ríos-Tamayo & Lyle, 2020: 295, f. 21A-J, 22A-C (mf)

Lepthercus sofiae Ríos-Tamayo & Lyle, 2020 | | South Africa urn:lsid:nmbe.ch:spidersp:051821

Lepthercus sofiae Ríos-Tamayo & Lyle, 2020: 298, f. 23A-J (Dm) Original description